ORNAMENTAL PLANTS IN THEIR NATURAL HABITATS
Rich floras are typical of all of the above regions, but the
Caucasus clearly displays the most diversity. The Caucasian mountains
lie between the Black and Caspian seas to the north of Turkey and Iran.
There are two main ranges, the Greater Caucasus and the Lesser Caucasus,
which run parallel from the northwest to the southeast for over 1,500
km/900 miles. Hollis and Brummit (1992) divide the Caucasus into two
parts: the North Caucasus, and the Transcaucasus. The former consists
of the northern slopes and ranges of the Greater Caucasus
and includes a number of members of the Russian Federation (former
autonomous republics and regions of the FSU: Kabardino-Balkaria, North
Ossetia, Checheno-Ingushetia, Dagestan, Adygeya, and Karachaevo-Cherkessya).
The Transcaucasus comprises the southern ranges of the Greater Caucasus
and all the Lesser Caucasus, and includes the Republics of Georgia,
Azerbaijan, and Armenia. A short botanical guide for travelers in these
three Republics are given below. A short description for the vicinity
of the Sukhumi, the Lake Ritza environs, mt. Mtirala, Georgian Military
road, Lagodekhi reserve; Zakataly reserve, Lenkoran’ and Talysh regions;
Garni, Gekhard, the Lake Sevan environs, mt. Aragats; for the Russian
Federation: Teberda reserve, Kabardino-Balkaria.
The Caucasus is mentioned in the Bible, when the waters of the Flood
subsided the Ark landed on Mount Ararat which served as a resting place
for Noah. The ancient state of Chaldea (called "Urartu" by the Assyrians),
was located in southwestern Asia, on the Euphrates River and the Persian
Gulf; and in the 9th century BC south Armenia was part of this country.
It was famous for its advanced agriculture, particularly known for its
diversity of wild wheats. To this day the same area is famous for the
excellent bread called Armenian bread, which stays fresh for a long period,
and no place on earth that can match the Transcaucasus in the number of
wild wheat or Triticum species. Here, in Georgia and Armenia there
are 8 endemic species of Triticum. These territories were the first
ones where the wheats were cultivated (Zhukovsky, 1964).
Vegetation of the Caucasus is remarkably luxuriant and rich. A large
area of the mountains is covered by forests, and near the snowline at ca.
2,700 m (9,000 ft), a magnificent alpine vegetation thrives. Snow-capped
peaks create spectacular landscapes, the principal peaks being El’brus
(5,633 m/18,468 ft), the highest mountain in Europe, and Kazbek
(5,047 m/16,558 ft). The southwestern Caucasus is characterized by a
warm and humid climate, but to the north and east this is more continental,
and in some places even arid conditions exist.
Various authors give different numbers for ornamental species in the
Caucasus. Grossheim (1952) estimated that there were about 5,000 species
in the Caucasus, of which 600 are ornamental. Other Caucasian botanists,
have suggested that more then 1,000 ornamental species are present. The
daisy family (Asteraceae) is prevalent, comprising one-eighth of
all the indigenous species. More than 800 species assort to 130 genera,
and this figure includes at least 40 genera with ornamental potential.
Also well-represented in the Caucasus are the buttercup (Ranunculaceae),
lily (Liliaceae), mint (Lamiaceae), pea (Fabaceae),
and carrot (Apiaceae) families. Over 60 families with ornamental
species have been recorded. Bellflower (Campanula) is one of its
genera richest in ornamental species, with ca. 40 species endemicaly
restricted to the Caucasus. Rich endemism is also shown by the iris
(Iris), cornflower (Centaurea) and primrose (Primula)
genera. The Caucasus has 17 endemic genera, found here and nowhere else
in the world e.g., Agasyllis, Chymsydia and Lepidotrichum of
carrot family, Sredinskya of primrose family, Woronowia of
rose family (Gagnidze et al., 2002).
A good rule of thumb is to seek new ornamentals in genera with species
already in garden cultivation, but the Caucasian flora amazingly contains
ornamental species that belong to genera totaly unknown in cultivation.
Recent discoveries in the daisy family include a very good groundcover
plant Cladochaeta endemic Caucasian genus with two species and a
beautiful pink-flowered perennial
Kemulariella with six species on the Caucasus. New species of
"old" genera are essential for hybridization and can play important roles.
For example, "new blood" is highly desirable for future paeony breeding
because most of the possibilities within Paeonia lactiflora seems
to be exhausted. Of special significance are the yellow-flowered paeonies
steveniana, P. tomentosa and P. wittmanniana. The crimson-flowered
P. caucasica may also be important for breeding purposes.
Caucasian lilies can also give many excellent hybrids. Of the six
Caucasian species, the most showy are a white-flowered
Lilium ledebourii and the lemon-flowered L. kesselringianum.
Iris, Gladiolus and Helleborus species may likely prove
valuble for producing hybrids. Species of the last genus or Christmas rose
have given rise to extremely diverse, natural hybrids whenever several species
grow together, as in Abkhazia. There, both white and richly dark red-flowered
forms have been collected. Large-flowered or curiously coloured forms are
seen restricted to certain areas. A white-flowered form of Stachys
macrantha, for example, grows near Bakuriani, and comes from seed in
cultivation. An impressively blue large-flowered Delphinium speciosum
(to 3 cm/ 1.2") has been collected throughout the entire range, and yet
another elegant small-flowered form of the same species occurs in the
northern Caucasus and Ossetia. Certain species exhibit unusual features
in different geographic areas, for example, sometimes Potentilla divina
can produce rather large pink flowers and white-hairy, ternate leaves,
Hylotelephium caucasicum develops creeping stems and deep dark red
inflorescences, and Caltha polypetala forms are seen with double
flowers with 7 to 10 sepals.
Extensive altitudinal distribution is characteristic of Caucasian species
having widespead ranges. However, some are endemic species and occur in
only one peculiar mountain zone. These species will require special
environmental conditions in cultivation. Ornamental Caucasian plants that
demonstrate very diverse habitat preferences are particularly valuable
because they adapt well to diverse enviroments and can be grown in different