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BFNA Title: Gollania |
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XX. GOLLANIA Brotherus in A. Engler and K. Prantl, Nat. Pfl. 1(3): 1054, figs. 755, 756. 1908 * [For William Gollan, d. 1905] Masanobu Higuchi Elmeriobryum Brotherus in
A. Engler and K. Prantl,
Nat. Pfl. ed. 2, 11: 204 Plants with dorsal, lateral, and ventral stem
leaves often different in shape and size. Stem leaves straight to
falcate, ovate-, oblong- or narrowly lanceolate;
margins recurved at base, serrate or serrulate, rarely entire distally, entire proximally;
costae double; median laminal cells linear, thin-
to thick-walled, smooth, rarely porose; alar cells differentiated, subquadrate
to rectangular. Branch leaves smaller and narrower than stem leaves. Sexual
condition dioicous; inner perichaetial
leaves oblong-lanceolate, filiform-acuminate, serrate, usually reflexed
at apices. Seta long, smooth, usually sinistrosely
twisted in proximal half and dextrorsely twisted
distally when dry. Capsule inclined to horizontal, ovoid to cylindric;
exothecial cells elongate-rectangular to subquadrate, rarely polygonal; operculum conic, sometimes
with a short beak; peristome double, exostome teeth
transversely smooth to striolate proximally,
papillose distally on the external surface, papillose and with projecting trabeculae on the internal surface; segments of endostome
teeth keeled, not split; cilia 2--4; basal membrane about 1/2 as high as
exostome teeth. Spores finely papillose. Species 20 (1
in the flora): nw North America; Asia ( SELECTED
REFERENCES Ando, H. 1957. The first record of Gollania
in 1. Gollania turgens (Müller Hal.)
Ando, Bot. Mag. Tokyo 79:
769, fig. 6 . 1966 Cupressina turgens Müller Hal.,
Nouvo Giorn. Bot. Ital. n. ser. 5: 196, 1896; Hypnum
turgens (Müller Hal.)
Stem to 7 cm, densely pinnate-branched,
rarely loosely branched; leafy stems complanate,
with secund leaves, hypnaceous
in appearance. Stem leaves well differentiated; dorsal leaves strongly
falcate, narrowly lanceolate, gradually narrowed to
a slender and long apex, subcordate at base,
plicate, 2.7--2.9 x 0.60--0.75 mm; margins markedly recurved
both at base of apex and below, irregularly serrulate
above, subentire proximally; costae double and
parallel, 1/4--1/3 the leaf length; median laminal
cells linear, weakly flexuose, lumen 60--80 x 2--3
µm, thick-walled, porose, smooth; alar cells well-differentiated, 6--8 in marginal row,
8--10 in transverse row; ventral leaves flat at apices. Branch leaves
narrower, 1.6--2 x 0.35--0.45 mm. Innner perichaetial leaves slightly reflexed
at apices; costae double or indistinct; laminal
cells thick-walled. [Seta 2.5--3 cm. Capsule horizontal,
cylindric, weakly arched, 2.2--2.8 mm exclusive of the operculum, 0.9--1.2 mm
wide; exothecial cells subquadrate,
rarely polygonal, 17--48 x 17--27 µm, thick-walled. Spores 18--22 \um.] Capsules
mature fall. Boreal forest zone to arctic tundra, mostly growing on wet or
moist rocks, often calcareous; 710--4730 m; B.C., Yukon; Alaska; Asia
(China, Japan, Nepal, Russia). Gollania turgens is known from 10 or fewer localities from North America and sporophytes not known from |
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