BFNA Title: Anacamptodon
Author: Piers Majestyk
Date: November 7, 2002
Edit Level: R Brum+
Version: 1

Bryophyte Flora of North America, Provisional Publication
Buffalo Museum of Science
1020 Humboldt Parkway
Buffalo, NY 14211 USA


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xxx. ANACAMPTODON Bridel, Musc. Recent. Suppl. 4: 136. 1819 [1818] • Greek anacamptos, reflexed, and odus, tooth, referring to the reflexed exostome teeth

Piers Majestyk

            Plants small to medium sized, in dull to shiny, yellow to green mats; stems creeping, somewhat radiculose, freely branched; branches ± ascending; leaves of stem and branches similar, erect to slightly spreading when dry, spreading to subsecund when moist, ovate to oblong-lanceolate, margins erect and entire to ± crenulate distally, costa single, ending at or beyond midleaf, cells smooth, ± rectangular and bulging at base, middle and distal cells short-rhomboid to linear. Sexual condition autoicous; perigonial leaves ovate, shortly acute, ecostate; perichaetial leaves oblong-lanceolate, long-acuminate, serrulate distally; costa faint, 1/2--3/4 the leaf length; cells elongate. Seta elongate, straight, smooth, yellow, brown or red. Capsules oblong-cylindric with an apophysis; strongly constricted below the mouth when dry; exostome teeth triangular-lanceolate, attenuate, thin, the teeth 16, paired, strongly recurved when dry, minutely papillose on both surfaces throughout; endostome segments 8 or 16, not arising from a basal membrane, filiform, elongate, 1/3--3/4 length of exostome. Operculum conic, obliquely rostrate. Calyptra cucullate, smooth, naked. Spores spherical, finely papillose.

Species ca. 12 (1 in the flora); temperate and tropical areas, North America, South America, Europe, Africa, Asia, Australia.

SELECTED REFERENCES: Crum, H. A. 1958. Anacamptodon in North America. Bryologist 61: 136--140. Crum, H. A. and L. E. Anderson. 1981. Mosses of Eastern North America. 2 vols. New York.

1. Anacamptodon splachnoides (Bridel) Bridel, Musc. Recent. Suppl. 4: 136. 1819 [1818]

Orthotrichum splachnoides, Musc. Recent. Suppl. 2: 4. 1812; Campylodontium hypnoides Schwägrichen; Anacamptodon splachnoides var. americanus Hampe

            Plants yellow-green to green; branches straight to slightly curved at tip; stem and branch leaves 0.7--1.4 mm, erect to spreading; costa 1/2--4/5 (occasionally shorter) leaf length; median and distal cells 30-56 ´  8-11 µm.  Seta 7--14 mm, yellow. Capsule 0.8--2 mm, yellow to brown, exostome yellow to light brown; endostome light orange to golden, 1/3--1 length of exostome. Calyptra 1.7--2 mm. Spores 10--15 µm.

            Mature capsules present Jan.--Dec. (mainly Apr.--Oct.). Bark of trees, wet, sheltered niches such as crotches, fissures or knotholes or rarely on logs, stumps or wet rocks; 0--1700 m; N.S., Ont., Que.; Ark., Conn., Fla., Ga., Ill., Ind., Ky., La., Maine, Md., Mass., Mich., Miss., N.H., N.J., N.Y., N.C., Ohio, Pa., Tenn., Tex., Vt., Va., W.Va., Wis.; Europe.

            The habit of Anacamptodon splachnoides combined with frequent sporophyte production with distinctive, strongly reflexed exostome teeth makes this one of the more easily recognized mosses. The most common substrates for A. splachnoides are knotholes or wet crevices of Fagus grandifolia, Betula spp. and Acer spp.