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BFNA Title: Conardia |
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Conardia - Amblystegiaceae CONARDIA H. Robinson, Phytologia 33: 294. 1976 * Howard A. Crum Lars Hedenäs Plants small, occasionally minute, light green or
yellow. Stems irregularly branched
± in one plane, with central strand and without a hyalodermis; paraphyllia
absent; rhizoids or rhizoid initials on stem and on back of leaf costa or on
lamina near leaf apex, warty-papillose, strongly branched (often
forming tomentum); axillary hairs with 1--4 distal hyaline cells. Stem leaves erect to spreading,
straight or slightly falcate, lanceolate or ovate-lanceolate, gradually
narrowed upwards, apex acuminate, not plicate, slightly concave; marginal
lamina cells unistratose, margin plane or partly slightly incurved or
recurved, finely, or near base often coarsely denticulate; costa single,
ending in acumen, sometimes percurrent; median lamina cells linear or
short-linear, smooth; differentiated alar cells usually numerous, quadrate or
more often rectangular, not inflated, forming a distinct transversely
triangular group, reaching from leaf margin to leaf middle at insertion or
almost so, decurrent. Sexual condition
autoicous (dioicous in Europe); inner perichaetial leaves ± suddenly narrowed
to acuminate apex, not plicate, margin irregularly denticulate above, lamina
cells smooth. Capsule cylindrical,
slightly curved, inclined; peristome slightly reduced; exostome margin
indistinctly dentate above. Spores
14.0--18.5 \um. Species 1 (1
in the flora): temperate North America, Central America, Greenland, Conardia has the appearance of a small Amblystegium,
but can be distinguished by its warty-papillose, much branched rhizoids that
are frequently inserted on the back of the costa or near the leaf apex.
Rhizoids can mostly be found in the basal portions of at least some shoots.
In addition, the distal parts of the axillary hairs consist of 1--4 cells
that are shorter and broader rather than the 1--2 narrow and delicate ones found
in Amblystegium. The capsule is only
slightly arcuate rather than curved throughout as in Amblystegium species. SELECTED
REFERENCE Hedenäs, L.
1989. On the taxonomic position of Conardia
Robins. J. Bryol. 15: 779--783. 1. Conardia compacta
(Müller Hal.) H. Robinson, Phytologia 33: 295. 1976 Hypnum compactum Müller Hal., Syn.
Musc. Frond. 2: 408. 1851;
Amblystegium americanum Grout; A. compactum (Müller Hal.)
Austin; A. dissitifolium
Macoun & Kindberg; A. holzingeri (Grout) Grout; A.
subcompactum Macoun & Kindberg; Brachythecium collinum var.
holzingeri Grout; Rhynchostegiella compacta (Müller Hal.)
Loeske; R. compacta subsp. Plants in dense, more or less loose mats. Stems
erect-ascending or creeping; pseudoparaphyllia foliose or filamentose. Stem leaves usually crowded,
often homomallous when dry, 0.6--1.2 mm, decurrent; costa commonly bearing
cylindric, papillose gemmae or rhizoids abaxially; laminal cells, 5--9:1,
shorter and broader at base and quadrate or rectangular in alar groups. Seta
8--14 mm, orange-yellow or at base reddish. Capsule annulus of 1-2 rows
of separating cells; operculum conical to shortly rostrate; exostome teeth
lanceolate, brownish yellow, at base reticulate or reticulate-striate or with
portions cross-striolate; endostome cilia mostly 1—2, up to half as long as
segments. Spores finely papillose. Damp cliffs,
especially limestone, swamps on logs, stumps, humus, bark at base of trees; low
to high elevantion; Greenland; Alta., B.C., Man., N.S., Ont., Que., Sask.,
Nfld., N.W.T., Nunavut, Yukon; Alaska, Ariz., Ark., Calif., Colo., Idaho,
Ind., Iowa, Kans., Mich., Minn., Mo., Mont., Nebr., Nev., N.Mex., N.Y.,
N.Dak., Ohio, Penn., S.Dak., Tex., Utah., Wash., W.Va., Wis., Va.; Mexico
(Baja California, Puebla); Central America (Guatemala);. n, c Europe; Asia
(Kashmir); |