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Data Set
from “Genera of the Pottiaceae: Mosses of Harsh Environments” by R. H.
Zander Bull.
Buffalo Soc. Nat. Sci. 32, 1993 |
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This data set contains morphological
information on the 76 genera of the Pottiaceae, and on ten (fully
capitalized) non-pottiaceous genera used as outgroups in analysis. A dash
(“-”) indicates a character that is variable in state or
otherwise “unknown, undefined, or missing” (Farris 1988). There
are 86 taxa and 75 characters; the first character is numbered as character
“0”. 0
1
2
3
4
5
6
7 012345678901234567890123456789012345678901234567890123456789012345678901234 POLYTRICHUM 1101100010120012000102103110111201000000-00--10-----0001012101110-----12020 BRYOBARTRAMIA 100-000100-0002000000110100-01-10-0000000011110-----101000100111-0----01102 CERATODON 11010011-011113100010111111011110100-00-000--00-----001-0121011-11110-0--00 DIPHYSCIUM 10000001002-002-0-0001-0201011120000000--0---10------0101121011-11-20-01021 DITRICHUM 1-01100-000-0-2-0-010--1-110111100000100000--00------0-1012101111112-101000 ENCALYPTA 0-0100--002-00-0000000-0200-11-20-00-001001111-01010-00-2-210111-1120-----2 GRIMMIA 110-00010011-1-00-0101-1000-00010-000000-00----01110-0--1-210111---20-0100- PTYCHOMITIUM 0101-1-000120022010111101010111100000000100--10-----10000121011111120-01--1 SYRRHOPODON 1100000-001---2-0--10-1-1010---10000000-00--11100110100-1121011101020--1000 TIMMIA 1101001-101200-200010210301011120100000200-1110------0010121011111-20-02-20 Acaulon 000000-1001-002-000100-1000-01110-100-010--0-0000000-00010000111------00121 Aloina -00-000-00-2001-00-012-0310-11120101200-110--1000000-00-0121011111121101020 Aloinella 1101001000-0001-00-01000210-11-1010100010--111000000001-0-21011111-20-01100 Anoectangium 11-110-1101-11-000010-110-0-0-110000000--0--1100000001110021011110----01000 Aschisma -00100100010002-00-100100010-0010-00000-001112000000100010100100------10100 Barbula 1-011--1101--1--000-0-110010-1110-00000100--1111101100--00210111-1221101000 Bellibarbula 11011011001-0130000101--00100111000000--001-1000000000100021011110----01120 Bryoceuthospora 000-0001101-002200010001101000010100000100100-000000-01010101011-------0101 Bryoerythrophyllum 1-0111-1101-----00010--0101011-10-000000-01111101010-0--0-2101111--2-101020 Calymperastrum 10-11101102-0020010102101010110101000001001121000000---------------------0- Calyptopogon 110011-11012--2-00110-11000-110101000011001-12100101-01001210111112211110-0 Chenia 100110111-210-3-000-0---000-011101000001000--1110000000-----01111-221101-2- Chionoloma 110101--100200-100110110301111020000000-001112000000---------------------0- Crossidium -00100-100-000--000--1-0100-01-1010110--0--111000000-0--0-21011111121101000 Crumia 110-001110220030011-00-1100-011100001001001-21000000000-0121011111121101-10 Dialytrichia 11011011101-1130010-0-11011011-101000001001111000000000-0121011101221101000 Didymodon 11-1101110---03-0-0-0---0----111000000-1-01-1-11111000--0-210111-1---1010-0 Dolotortula 11010001112-0030011000-1000-011101002001000--0000000000101210111112211--02- Erythrophyllastrum 1101-1111011-0200-0102103010011101000001-0111100000000110121011111020-01020 Erythrophyllopsis 11-101111012-020000102102110111101000001101111000000000-0121011111-20---02- Eucladium 1-00-10110-1002110010-1-101011-1000000-1001-1100000000001021011101120-01000 Ganguleea 10-0-00011210010000000-0000-011100000002000--100000011100021011110------00- Gertrudiella 110111011012003-000101-0300-11121100000200-0110000000--------------------1- Globulinella -0010--100-000-00000100010--0111011000--000--110000000000-21011111120-01000 Gymnostomiella 1001-0111020002-000-00-1000-00110000100100101110111000100021011100----010-0 Gymnostomum 11-10-11101000-000010-0-0010-1-10000000-001-11100000000000210111-0----0-000 Gyroweisia 100-101100-00--0000-0--110--01-10000000--01-11110000-0-00-2101111-020-0-0-0 Hennediella 1-01---110--002-01110-1-100-01-101002000001-21000000-00-2-2101111--2110--20 Hilpertia 1-01-0-1001-003-01010021000-010101002--101-01011000010100-21011111221101-20 Hymenostyliella 110110100012001-0001001-1010111100000012000--100000001100-21011100----01000 Hymenostylium 11-0101110--11-000010-11001000-100000--000101100000000000021011100----00000 Hyophila 1--11-001-2100--000-00-0101011110-00000-00-011101011-0000-21011110----01-00 Hypodontium 11010110101-00---01-0210201-11020000000200110200000000102121011101100-11100 Leptobarbula 1001-01110100020000100-0011011110000000000111100000000100-21011111221101000 Leptodontiella 1100111110111131000101-11110100100000000001-1-1010100010002101101111000110- Leptodontium 11001--1101-113-00010--111101001000000--001-11101110-01-0121011-111-0-0-100 Luisierella 000000101---00-0000-01001-0-100100000002000--20000001000012101111--20-010-0 Microbryum 000--0-1002-003-000100--100-011101001101001-1-000000100020--01110--20--0120 Mironia 11011011101-113-0101021-111011110100000000111110000000110-21011111221101020 Molendoa 11-11-0-10--00----010---1-1011110000000-00111110101001100021011100----01000 Pachyneuropsis 110100111002002001010210301-111200000001001112000000---------------------0- Phascopsis 10010-1100-00020000-001-100-01010-000001001-11100000000010-00111------00122 Plaubelia 10011010112-001-000-00-010-001110100000200-00-10100000-00121011111120-0-000 Pleurochaete 110111011012002100110-102-1011-10000000-001112000000011101210111112211020-0 Pseudocrossidium 1-01--01101---30000-0--011-011110101000100111110111000--0121011111-211020-0 Pseudosymblepharis 1101-1-110--00-000010210301111-20000000-00111200000000--012101111--20-01000 Pterygoneurum -00100-1001-001-000---20100-01110100000-00-01-00000010002-2101110-020-0-10- Quaesticula 10010000101100100000-000101001110000000-001001000000000-0021011111221101000 Reimersia 011010-1101-1120000102110010101100000-10000--100000000000021011100----01000 Rhexophyllum 110111-11011113200010211011010110-00000110111100000000110121011110----01020 Sagenotortula 11010001002-0020000100--000-110101003001000--111000000010-21011111221102020 Saitoella 1001000100-000300110100-100-01-0-0100001001111000000---------------------2- Sarconeurum 1-0-0-11101-002000010-1000-0010101001000001-11100000---------------------2- Scopelophila 110000-11021-1--00010---000---110000000-000--110101000000-210111-0----00000 Stegonia 00010011001100--000-00-1100-01-101101101010--1000000100-0---01111--20-0-100 Stonea 000000110020002-000110-0000-01-1001000010010111000000--------------------2- Streptocalypta 10010--110--002-00-----010-01102-000000-001112000000-000002101111-221101000 Streptopogon 1-0010-1001--03-0------1000-0--1000021-1000--0100100-0-0012101110--211-1--1 Streptotrichum 1100111110121132000102-1111010010000000000102100000000101121011011120-01100 Syntrichia 110-0--1102-----0-0----0000--101--0000-1-01-11110100-0012-2101111122110-020 Teniolophora 11011010102100--011-0010201011110000000200-001101010000------------------0- Tetracoscinodon 110010-10001112000010-1-1010111100000001001111000000001100210111111-0-01100 Tetrapterum 1001001110110020000101102-1011110100000-001111000000101011200101------011-0 Timmiella 1-0111-011120022000102103-10111201000002100--1000000-0010-210111-1---002100 Tortella 110-0101101-00200001011-101011-10000000-001112100000-01-012-0111--22110--00 Tortula 1-01001110--0-3-0---0---100-01-101100001001-11000000-00-2---01111--2110--00 Trachycarpidium -001001110110-20000100-1101011-10-00100-001112000000100010201111------00101 Trachyodontium 110011111012113201110211111010010000000-00102100000000110221011011120----0- Trichostomum 11---1--101-002-0-01-1101-10-1-100000001001111000000-00-0-2-01111---0-01--0 Triquetrella 111-10111011113-000101110010-011000000--001-00000000001101210111111-0-00000 Tuerckheimia 11-100--10-1002100010-10101011-1000000-1001-2-000000000000210111--0-----00- Uleobryum -001000110-0012000--001110100-0101000001001-11000000100010101011------10101 Weisiopsis 1001-01010-10-20000-01--0-0-0-11000000-2000--1000000100000210111--1-0--1000 Weissia 100101101011000000010-10101011-10-00000-00111-000000-0--0---01111---0-0-100 Weissiodicranum 110100-00011000000010-1010101111000000020011-1000000-00------------------0- Willia 11010011102-012-00010---100--1010100000100111100000000100-2101111-22110-02- |
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The character states scored for the data set are listed
below; (R) = reduction-related character, of which there are 22 and weighted
lower than other characters in most cladograms, (0´) = character not used at all in Cladogram 15 (i.e.,
not phylogenetically informative), (1´) = character occurred once in Cladogram 15 (i.e.
phylogenetically very informative), and * = the hypothesized plesiotypic
character state for the Pottiaceae. Character numbers correspond to those in
the data matrix used with Hennig86, and begin with zero. 0. (R, 0´) plant conflux 0. gregarious or scattered 1. *caespitose, usually in
a mat or turf 1. (R) length of
stem 0. short, less than 1.0 cm,
usually less than 0.6 cm 1. *relatively long,
usually 1.0 cm or greater 2. (1´) stem transverse section shape 0. *rounded-pentagonal 1. triangular 3. stem central
strand presence 0. absent 1. *present 4. stem
sclerodermis presence 0. *not or little
differentiated from central cylinder cells 1. clearly differentiated 5. stem
hyalodermis 0. *absent 1. present (sometimes small) 6. (1´) axillary hair basal cell walls 0. hyaline and all cells of
hair similar 1. *of 1 or more cells with
thicker or darker colored walls 7. leaves when
dry 0. tubulose 1. *occasionally channeled
but not distinctly tubulose 8. leaves when wet 0. appressed to weakly
spreading 1. *widely spreading to
squarrose 9. (0´) leaves, size gradation 0. *gradually becoming
larger distally on stem 1. rosulate (abruptly
larger distally) 10. leaf shape 0. long-triangular to
linear-lanceolate 1. *lanceolate 2. broadly
ligulate-elliptical to spathulate 11. (R) leaf
length 0. less than 1.5 µm 1. 1.5 µm to 3.0
µm 2. *3.0 µm or more 12. leaf ventral
surface above midleaf 0. *nearly plane to broadly
channeled 1. keeled 13. rather deep,
narrow groove along costa 0. *absent 1. present 14. margins 0. sharply incurved 1. *incurved or involute 2. plane 3. recurved to revolute 15. marginal
toothing 0. entire or minutely and
evenly crenulate 1. denticulate or serrulate 2. *strongly toothed 16. margins
specially denticulate 0. *not denticulate only
below 1. denticulate only near
leaf base or at top of leaf sheath 17. (1´) upper marginal cells differentiation 0. *same number of layers
as medial cells 1. always differentiated as
a bistratose (or more) border 18. upper
marginal cells, elongation 0. *not longer than medial
cells (sometimes larger) 1. rectangular and clearly
longer than medial cells 19. (1´) leaf apex 0. rounded 1. *acute to acuminate 20. leaf apex
ventral surface 0. *flattened, channeled or
keeled 1. cucullate 21. (R) leaf
base 0. little differentiated in
shape 1. ovate to elliptical 2. *sheathing the stem and
commonly oblong and with “shoulders” 22. (1´) costa 0. ending before the leaf
apex 1. *percurrent or apiculate
or short-excurrent as a mucro 2. excurrent as an awn 23. costal
ventral cells viewed from above 0. *quadrate to very
short-rectangular, occasionally in many layers 1. longitudinally elongate
3:1 or more 24. number of
rows of cells across ventral surface of costa 0. usually 2 but up to 4 1. 4–6 2. 6–10 3. *usually 10 or more 25. shape of
transverse section of costa 0. *round to semicircular 1. distinctly flattened,
usually reniform 26. ventral
stereid band 0. absent 1. *present 27. (1´) size of ventral stereid band 0. *smaller than the dorsal
or of nearly equal size 1. larger than the dorsal 28. transverse
section of dorsal stereid band 0. round or semicircular 1. *clearly flattened or
ventrally indented, reniform or crescent-shaped 29. ventral
costal epidermis 0. absent 1. *present 30. dorsal
costal epidermis 0. absent 1. *present 31. guide cells 0. usually 0 1. 2–6 2. *commonly more than 6 32. (0´) number of guide cell layers 0. *1–2 1. 3 or more 33. hydroid
strand 0. absent 1. *present 34. (R, 0´) ventral costal outgrowths 0. *absent or not a bulging
pad of cells 1. a bulging pad of cells 35. (R, 1´) ventral costal filamentous outgrowths (note:
exclusive of lamellae) 0. *absent or not of
filaments 1. of separate filaments
three or more cells in length 36. width of
medial upper laminal cells 0. *small to medium sized,
7–14 µm 1. rather large,
15–17 µm 2. very large, 18–25
µm 3. extremely large, 25
µm or more 37. (0´) length to width ratio of medial upper laminal cells 0. *1–2:1 1. 3–4:1 or more 38. (0´) upper laminal cell walls 0. *thin to evenly
thickened, lumens quadrate to rounded 1. trigonous or porose,
lumens irregularly angular or stellate 39. superficial
walls of upper laminal cells 0. flat or very weakly
convex on both sides 1. strongly convex to
bulging on both sides 2. *ventrally
bulging-mamillose, weakly convex dorsally 40. layering of
upper laminal cells 0. *medially unistratose 1. medially bistratose, at
least in patches 41. (R, 0´) dorsal superficial laminal cell walls 0. *about same thickness as
the ventral or weakly thicker near costa 1. throughout distinctly
thicker than the ventral 42. (1´) upper laminal papillae 0. absent 1. *present 43. kind of
papillae when present 0. simple, hemispherical 1. *bifid to multifid to
columniform to capitulate 44. number of
papillae per lumen when present 0. one or occasionally two 1. *2–6, usually
bifid or multifid 2. many, usually 6 or more,
simple or bifid 45. basal cell
group 0. not or little
differentiated from upper medial cells 1. *clearly differentiated
approximately straight across leaf or rising higher medially 2. differentiated as a vee,
with at least laterally differentiated cells rising higher marginally as a
border tapering distally 46. propagula
presence 0. *always absent or
extremely rare and associated with unusually moist conditions 1. often present and
characteristic 47. (1´) position of propagula when present 0. not borne on rhizoids as
brood bodies, or absent 1. borne on rhizoids as
brood bodies 48. position of
propagula when present 0. not born in leaf axils,
or absent 1. borne in axils 49. (1´) position of propagula when present 0. not borne on leaf costa
or lamina, or absent 1. born on leaf costa or
lamina 50. shape of
axillary propagula when present 0. not clavate or
filamentous 1. clavate or filamentous 51. shape of
axillary propagula when present 0. not branching or
stellate 1. branching or stellate 52. (R) sexual
condition 0. *dioicous 1. monoicous 53. (0´) perichaetium 0. *terminal on main axis 1. lateral on main axis at
ends of very short branches 54. perichaetial
leaves 0. *similar to cauline
leaves or occasionally smaller or somewhat enlarged 1. distinctly different in
size or morphology, sometimes strongly sheathing 55. (R) length
of seta 0. nearly absent to short,
less than 1 cm 1. *elongate, 1 cm or
longer 56. (R) seta
twist 0. *usually twisted
clockwise 1. never twisted 2. usually twisted
counterclockwise 57. (R) theca
length 0. short, 1.5 µm in
length or less 1. *usually between 1.5 and
3.5 µm in length 2. 3.5 µm in length
or more 58. (R, 1´) shape of theca 0. spherical and
non-apiculate 1. spherical and apiculate 2. *ovoid to cylindrical 59. (R, 1´) capsule dehiscence 0. cleistocarpous 1. *stegocarpous 60. (R, 1´) capsule ornamentation 0. *surface nearly smooth 1. surface evenly mamillose
or with distinct protuberances of strongly bulging cells basally or
throughout 61. (R, 1´) exothecial cells 0. with walls superficially
thickened centrally and lens-like 1. *with evenly thickened
superficial walls 62. (R, 0´) capsule 0. cleistocarpous and
rupturing mainly along weak transverse walls at butt ends of long-rectangular
exothecial cells 1. *rupturing irregularly
if cleistocarpous, or stegocarpous 63. (1´) stomates 0. absent 1. *present 64. annulus 0. of weakly differentiated
cells 1. *of vesiculose cells,
often in two or more rows 65. (R)
peristome teeth 0. absent 1. *present 66. (R)
peristome teeth when present 0. rudimentary, a low plate
or short-elliptical in shape 1. 16, often cleft to near
base into two or more rami, or of 16 pairs 2. *32 similar rami 67. (0´) ornamentation of peristome teeth 0. smooth 1. striate or ridged 2. *spiculose or papillose 68. (R)
peristome teeth twist 0. untwisted or very weakly
so 1. *distinctly twisted 69. (0´) peristome teeth if twisted 0. clockwise 1. *counterclockwise 70. (0´) calyptra ornamentation 0. *smooth or nearly so 1. papillose, distinctly
roughened or strongly mamillose 71. (R) length
of calyptra 0. short, less than 1
µm 1. 1–3 µm 2. *more than 3 µm 72. (R) spore
diameter 0. *8–15 µm 1. more than 15 µm 73. KOH color
reaction of upper laminal cell walls 0. essentially yellow 1. essentially orange 2. *essentially red,
usually a definite brick red 74. (R, 1´) calyptra shape 0. *cucullate or at least
long-conic and cleft, usually more than 1 µm in length 1. mitrate or short-conic
and uncleft, commonly lobed, usually less than 1 µm in length 2. campanulate-inflated |
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