Preliminary Cryptogamic (Moss, Lichen and Liverwort) Flora of the Canadian and American Gorge at Niagara Falls 1. Mosses

P. M. Eckel
Res Botanica, a Missouri Botanical Garden Web site

http://www.mobot.org/plantscience/ResBot/index.htm

June 21, 2004

 

PRELIMINARY CRYPTOGAMIC (MOSS, LICHEN AND LIVERWORT) FLORA OF THE CANADIAN AND AMERICAN GORGE AT NIAGARA FALLS

1. Mosses

 

P. M. Eckel, Missouri Botanical Garden, PO Box 299, St. Louis, MO 63166-0299 USA

 

 

 

Grimmia alpicola Hedw.

 

USA: Eckel below between steps and Whirlpool 422386 BUF

 

USA: [var. alpicola] Devil's Hole, near bottom with Bryum capillare var. flaccidum, Entodon seductrix, Bryhnia graminicolor, May 21, 1988, Eckel No. 9303181C (BUF). 

 

USA: Devil's Hole, upper slopes, Eckel 226386.

 

USA: (var. alpicola) Goat Island on the Niagara River, wooded slope just N of first 3-Sisters Islds Bridge, on dolomite boulder. Plants to over 7mm, lvs to 2mm, capsules immersed. P. M. Eckel 881003 Sept. 29, 1988 (BUF).

 

USA: (var. alpicola) Goat Island on the Niagara River, in depression on dolomite block of first bridge to the Three Sisters Islands. Plants to over 7mm, lvs to 2mm, capsules immersed. P. M. Eckel 881002 Sept. 29, 1988 (BUF).

USA: (As Schistidium alpicola). Luna Island, 1 Nov. 1988, Buck 16319 (NY).

USA: Green Island on the Niagara R just N of Goat Island. Old dolomite ballast on S side, occasionally submersed, wet to moist river margin. PM Eckel, Marguerite Eckel 9004158 April 15, 1990 (BUF).

USA: (As Schistidium alpicola): Second Sister Island, east end, 2 Nov. 1988, Buck 16411 (NY).

USA: var. alpicola. Dolomite bridge, First Sister Island,  in depression, with lichens, Eckel, Sept. 29, 1988 (BUF).

Grimmia alpicola var. rivularis (Brid.) Wahlenb.

 

CAN: Niagara Glen, just N of Whirlpool. Moist thin soil in depressions in limestone boulders. w. Tortella tortuosa, Didymodon rigidulus. With R.H.Zander. P.M.Eckel NF17 Feb. 19, 1983 (BUF).

 

Grimmia apocarpa Hedw. NY CHECK as var. apocarpa

 

USA: Zander, Devil's Hole lower area between Devil's and whirlp on USA side.

 

CAN: Grimmia apocarpa, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

As Grimmia apocarpa var. stricta, Drumm. Musc. Bor. Am. No. 55 by Macoun & Kindberg:] "... Niagara Falls (Macoun.)," Macoun & Kindberg, 1892, p. 62‑63. 

 

Grimmia calyptrata A doubtful record.

 

A western species collected by Drummond in the Niagara gorge in BUF is misplaced from collections he probably made out west. It is annotated as a new sp. Herb. No. 56168, . "On stones, Niagara, rare" annotated as Grimmia calyptrata Hook. on packet.  cf. Wils., Hook. J. Bot. This specimen is actually Grimmia calyptrate but misplaced, sec. Zander.

 

Grimmia pilifera P. Beauv.

 

2005 Rare Plant Bryophyte Status List  S3? [sic]

 

Grimmia pennsylvanica Schwaegr. ex Sull. & Lesq.

 

CAN: "near the Falls of Niagara; upon rocks (as Grimmia pennsylvanica), Drummond, Musci Americani No. 56 (NY). 

 

CAN: As Grimmia pennsylvanica, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN:  Niagara Glen (Foster's Flats) on the Niagara River. Dry, occasional seepage in dolomite caprock, upper path, with Didymodon rigidulus, Gymnostomum aeruginosum. With R. Zander. P. M. Eckel 2231185 April 5, 1985 (BUF).

 

Gymnostomum aeruginosum Sm. NY CHECK

 

Gymnostomum calcareum, Nees, Hornsch. & Sturm

 

Gymnostomum rupestre Schleich. ex Schwaegr.

 

[USA?] On rocks, Whirlpool, Niagara Falls. G. W. Clinton (with Seligeria sp., label indicates S. recurvata, but removed from packet) (NY).

 

CAN: As Gymnostomum rupestre Schleich. following Crum & Andeson p. 283, Queen Victoria Niagara Falls Park.  Cameron 1895

 

CAN: As Gymnostomum calcareum, Nees & Hornsch. [G. tortile, Drummond. Musc. Bor. Am. 23]

 

CAN: "... on calcareous rocks near the Whirlpool, Niagara Falls. (Macoun.)," Macoun & Kindberg, 1892, p. 12.

 

CAN: As Gymnostomum rupestre, Schwaegr., "... near the Whirlpool, Niagara Falls (Macoun.),"   Macoun & Kindberg, 1892 p. 13. 

 

CAN: Macoun & Kindberg (p. 13, 1892) reported (as G. rupestre) this species growing "near the Whirlpool, Niagara Falls ... (Macoun)."

 

CAN: As Gymnostomum calcareum NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: (as Gymnostomum rupestre) Hab. on ground towards 3 sister islands, Niagara Falls, Aug. 21, 1886, c. fr., Eugene A. Rau, Bethlehem, Pa. (NY)

 

USA: On Three Sisters Island, Niagara Falls, N.Y., Aug. 21, 1886, c. fr., AAAS meeting, Elizabeth Gertrude Britton (NY).

 

A specimen from NY determined as Barbula convoluta was identified by R. Zander (1996) to be Bryoerythrophyllum recurvirostrum mixed with Gymnostomum aeruginosum. The plants were collected at "Niagara Falls" by Wolle in 1873. There is a packet with Wolle's autograph on it, and the packet was constructed out of the fragment of a letter, presumably to Wolle, by Eugene A. Rau; the letter is unreadable due to its fragmentary nature.  Crum & Anderson (l981) discuss the occurrence of this species in North America and believe specimens identified as G. calcareum in eastern North America are in fact G. aeruginosum based on a variety of characters, p.  285. Zander, Bryologist 80:233‑69, 1977, synonymized G. calcareum with G.  aeruginosum.  Podpera (1954) reported a specimen of Gymnostomum calcareum from "Niagara." Macoun & Kindberg (p. 12, 1892) reported Gymnostomum calcareum "on calcareous rocks near the Whirlpool Niagara Falls.  (Macoun). 

 

USA: boulder field, halfway down Whirlpool steps, with Anomodon viticulosus, Bryum lisae var. cuspidatum, Tortella tortuosa, Campylium chrysophyllum, Encalypta procera, Plagiopus oederiana, Mnium thomsonii, Marchantia polymorpha, May 6, 1987, Eckel 9302092b (BUF); DeVeaux (Whirlpool) stone steps down calcareous gorge face, moist shaded wooded bottom above Niagara riverbank, on moist boulders, c. fr., with Bryoerythrophyllum recurvirostrum. Eckel Oct. 4, 1991 (BUF).

 

USA: Zander, Devil's Hole waterfall.

 

USA: Zander, Ongiara, lower slopes

 

USA: Zander, Prospect Park, lower slopes.

 

USA: At Niagara Falls, base of Goat Island, just below lower entrance to elevator, about 70 feet above waterline, in vug in large calcareous boulder. R. Zander, P. Eckel, W. Buck, R. Harris 8811012 Nov. 1, 1988 (BUF).

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Upper path, dolomite caprock, with Didymodon rigidulus, Grimmia pilifera. With R. Zander. P. M. Eckel 2131185 April 5, 1985 (BUF).

 

CAN: Above Queenston, on the escarpment just S of Brock's Monument on the gorge lip: Locust" Grove picnic area. Thin soil over moist rock" in stream bed. P. M. Eckel 232085 April 20, 1985 (BUF).

 

Haplocladium microphyllum (Hedw.) Broth. NY CHECK

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring). P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

CAN: (Welland Co., Bertie Twp.) Niagara River, Navy Island, 43*03'N, 79*01'W, just S of Niagara Falls; west side, bark base of tree; Oak woodland. Dicranum montanum, Ceratodon purpureus.  P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Hedwigia ciliata (Hedw.) P.‑Beauv. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron, 1895.

 

CAN: As Hedwigia albicans Lindb.  "... on rocks, Whirlpool Rapids, Niagara River, 1901. (Macoun.)," Macoun, 1902, p. 225. 

 

USA: Second Sister Island, west end, on glacial erratic, 2 Nov. 1988, Buck 16380 (NY).

 

CAN: (Welland Co., Bertie Twp.), Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. With Hypnum pallescens on granite glacial erratic; Fraxinus-Carya woods P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Helodium paludosum (Sull.) Aust. 

 

As Elodium paludosum Aust. "... Queenston and "roundhouse woods," near Niagara Falls, Ont., 1901. (Macoun.)," Macoun, 1902, p. 286. 

 

USA: Erie Co. Grand Island. Ubiquitous on Grand Island. Niagara R. Ann Glowny 1338 3-28-67 (BUF).

 

CAN: Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Forest interior; moist Red Maple forest. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Muck soil, with Climacium americanum P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

Herzogiella striatella (Brid.) Iwats.

 

Plagiothecium muehlenbeckii BSG.

 

CAN: "... on old logs at Foster's Flats, Niagara River, Ont., 1901. (Macoun.)," Macoun, 1902, p. 299. 

 

Homalotheciella cubcapillata (Hedw.) Broth.

 

2005 Rare Plant Bryophyte Status List. SH

 

As Homalothecium subcapillatum Schp. Devil's Hole, Niagara River. Day 1883.

 

Homomallium adnatum (Hedw.) Broth.  NY CHECK

 

USA: Devil's Hole, upper slopes, Eckel 726386  (BUF); (Eckel, l986b).

 

CAN: Niagara Glen (Foster's Flats). Boulder field. Base of dolomite block, shaded woodland, with Didymodon rigidulus Bryum capillare. P.M.Eckel 229486 April 11, 1985 (BUF)

USA: First Sister Island, western half, 1 Nov. 1988, Buck 16342 (NY).

USA: Second Sister Island, west end, on glacial erratic, 2 Nov. 1988, Buck 16364 (BUF, NY).

Hylocomium brevirostre (Brid.) Fleisch. NY CHECK

= Hypnum brevirostre (Brid.) Ehrh. 

 

Hylocomium splendens (Hedw.) BSG

 

Hypnum splendens Hedw.

 

CAN: As Hypnum splendens, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

Hymenostylium recurvirostrum (Hedw.) Dix.

 

= Gymnostomum recurvirostrum Hedw.

On August 8, 1865, George Clinton recorded in his journal that this moss was "everywhere common on wet rocks" below the Biddle Staircase (on Goat Island, west facing face of the Niagara River gorge)..

[USA?] Hab. Niagara Falls, July 18, 1874, Coe Finch Austin, leg. with Didymodon tophaceus (NY).

 

USA: Three Sisters Islands, Niagara River, Aug. 1886. L.M.U.  [Underwood] (NY) with bistratose leaf margins in patches. 

 

USA: Hab. ad cataractam Niagarae. Musci Alleghanienses, W. S. Sullivant, 1845, No. 193 (As Gymnostomum curvirostrum Hedw.), (NY).

 

USA: Rocks. Niagara Falls, N.Y., July, 1865, c.fr., G. W. Clinton, det. Charles H. Peck (NY).

 

USA: Biddle's Stairs, on damp rocks at Niagara Falls below Biddle's Stairs, Goat Island, Sept. 1853, c. fr., Chapman collection (NY)

 

USA: Hab. in rupibus irriguis Niagarae cataractae. Sullivant & Lesquereux, Musci Bor.‑Amer. ed 1, No. 41 (As Gymnostomum curvirostrum Hedw.)(NY). This collection has capsules which differ from the typical variety by the "capsula ovato‑cylindrica fusco‑lutea," but is otherwise indistinguishable.

 

USA: Hab. in rupibus irriguis Niagarae cataractae. Sullivant & Lesquereux, Musci Bor. ‑Amer. ed. 2, 1865, No. 53 (NY). The NY specimen from Jaeger's Herbarium (No. 777) was annotated as var.  ovatum Sull. & Lesq. It is still "capsula ovato‑cylindrica fusco‑lutea". The specimen examined above as No. 41 of the first edition represented a population of medium to small stature. In No 53, two specimens, one tall, relative to the other, are glued to the card, the lesser probably the var. inferior.  [also specimen NYS with two specimens on a card].

 

USA: Niagara Falls, G. W. Clinton ("3") several specimens glued to cards (NYS).

 

Canada. Hab. Falls of Niagara. Drummond, Musci Americani (Rocky Mts.) No. 25 (as Gymnostomum recurvirostrum (NY), a tall stemmed group and a small glued to the card, the smaller probably representing the var. minor. Crum and Anderson (1981) indicate that No. 26 of this exsiccat is var. minor. Two other examples seen of No. 25 (both NY) are single collections of one stature. Another specimen from the National Herbarium (CANM) in Ottawa "Falls of Niagara" No. 25. "Welland Co." in pencil.

 

USA: [?] Niagara Falls. legit G. W. Clinton (label of Charles H. Peck, Albany, NY) apparently distributed by Peck to Dr. E. C. Howe (NY). Two collections in fruit representing elongate‑stemmed population and short‑stemmed, both in fruit. Perhaps this particular mode of preserving specimens of Hymenostylium recurvirostrum was to demonstrate the variability of the species, as against naming, for example, short representatives var. inferior, or robust ones with cylindric capsules distinguished by Sullivant and Lesquereux.

 

USA: [?] Niagara Falls, Rev. Francis Wolle, 1873 (NY, in packet of Funaria hygrometrica).

 

[USA?] Niagara Falls, 26 August, 1917, c. fr. A. T. Beals (NY).

 

[USA?] Niagara. c. fr. 1873, Francis Wolle (NY). [ex herb. Eugene A. Rau, Bethlehem, Pa.]

 

[USA?] Niagara Falls. Aug. 21 & 22, 1886. c. fr. Eugene A. Rau, Bethlehem, Pa. (NY)

 

[USA?] Rocks. Niagara. North America. ex Herb. William Mitten (NY).

 

[USA?] On limestone. Niagara. Ruth Schoernhurst, July 17, 1937, c. fr. (NY).

 

CAN: Welland Co., limestone cliffs along Niagara River opposite American Falls, in seepage, Howard Crum no. 26, October 5, 1964 (CANM).

 

A specimen at COLO labeled "Falls Niagara C. W. July 1850", COLO label indicates New York with the possibility of the collector being T. P. James. Specimen mounted on card and hand labeled.

 

As Gymnostmum curvirostrum, "... at Niagara Falls. (Drummond.)," Macoun & Kindberg, 1892 p. 13. 

 

As Gymnostomum curvirostrum Hedw. Queen Victoria Niagara Falls Park.  Cameron 1895. 

 

As Gymnostomum curvirostrum Hedw. Niagara Falls, near the American staircase, Charles Peck in Day, 1883.

 

Podpera (1954) reported an occurrence from "Niagara." Macoun (p. 13, 1892) cited a Drummond specimen from Niagara Falls. Lesquereux & James (1884. pp.54): "Limestone rocks, and on deposits of carbonate of lime or tufa, near springs; very abundant at Niagara Falls."

 

USA:  on rock at base of American Falls, Prospect Point. W. Specht & J. McCandless Oct. 9, 1973 (BUF).

 

CAN: Gorge of the Niagara River, "First Gorge" road into gorge from Queenston, bottom slope, side of gorge, below Lewiston Bridge, wet clay near waterfall R. H. Zander 3723 Sept. 29, 1971 (BUF).

 

CAN: (As Gymnostomum recurvirostrum) Upper Great Gorge of Niagara River, below Table Rock House, near Horseshoe Falls. (spray zone) R. H. Zander Sept. 7, 1971 (BUF); Niagara Falls, near Horseshoe Falls, near bottom of slope, spray zone, on thin soil on rocks with grasses. R. H. Zander 3686b Sept. 7, 1971 (BUF).

 

CAN: Foster's Flats, (Niagara Glen) on the Niagara River, half way down. N-facing, moist, dolomite boulder. RH Zander 63585 Apr 5 1985 (BUF); Niagara Glen, half way up slope from river, Pot Hole Path, boulder, thin soil. R. H. Zander 3732a Sept. 29, 1971 (BUF).

 

USA: Gorge of Niagara R. at Devil's Hole, top of gorge, on limestone.

R Zander 4252a 12 Sept 1975 (BUF). 

 

CAN: Niagara Falls. Forming large tufaceous mass on limestone wall of gorge at the falls. I Herrnstadt & R Zander Mar 2 1977 (BUF).

 

USA.: Ongiara Gorge, between steps and Whirlpool, top of talus slope, base of dolomite caprock in seepage: frozen, with Didymodon tophaceus, northwest facing wall. P. M. Eckel 222386 March 21, l986 (BUF).

 

CAN: Niagara Glen (Foster's Flats). Boulder field. Base of dolomite block, shaded woodland. P.M.Eckel 525086 April 11, 1985 (BUF); Niagara Glen (Foster's Flats) on the Niagara River. Moist dolomite boulder, N-facing, half way up gorge, with Didymodon rigidulus and Desmatodon porteri. With R. Zander. P. M. Eckel 53585 April 5, 1985 (BUF).

 

USA: lower Arch Bridge, upper path USA: Eckel between steps and Whirlpool, seeps, path by caprock, Eckel 321386 (BUF).

 

USA: Niagara River Gorge, Sides Of Sheer Dolomite Cliff, Seepage: Abundant Bryophytes, Upper Old Rr Path Near Top Of Gorge By Schoelkopf Museum. P. M. Eckel 79386 Sept. 21, L985 (BUF).

USA: (as Hymenostylium recurvirostre) just outside spray area of Horseshoe Falls, boulder, thin soil, midslope, with Tortella fragilis, Hyophila involuta, Trichostomum crispulum, Zander 3484d, Oct. 28, 1979 (BUF), on rubble, talus slope, Zander 3491b, Oct. 28, 1979 (BUF); just outside spray area of Horseshoe Falls, vertical rock face of gorge, with Preissia quadrata, Zander 3492a, Oct.28, 1979 (BUF); spray area of Horseshoe Falls, near river, soil under boulder, Zander 3478, Oct.28, 1979 (BUF); spray zone of American Falls, talus, thin soil, crevices of rock, Zander 3444a, Oct. 28, 1979 (BUF); base of island, 1 Nov. 1988, Buck 16354 (BUF, NY).

CAN: Niagara Falls gorge, near the Horseshoe Falls along road to power station. Wet, dripping dolomite walls. R. H. Zander 3651 Aug. 10, 1971 (BUF); Niagara River Gorge, near Horseshoe Falls, down road to powerhouse, Lockport Dolomite, crevices R. H. Zander 3648b Aug. 10, 1971 (BUF).

USA: Luna Island, 1 Nov. 1988, Buck 16331 (NY).

Hyophila involuta (Hook.) Jaeg. & Sauerb. NY CHECK

 

2005 Rare Plant Bryophyte Status List S1

 

[No country] Hab. Niagara Falls, 1873, F. Wolle (NY).

 

USA: Zander, Devil's Hole, lower slopes.

 

USA: Gorge of Niagara River, riverside limestone boulders. R. H. Zander Sept. 12, 1975 (BUF).

 

CAN: Gorge of Niagara River at Niagara Glen, about half way between water and lip of gorge, side of moist, shady limestone boulder. R. H. Zander April 7, 1984 (BUF) .

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Moist dolomite boulder in solution cavity: the smaller plant is Gymnostomum aeruginosum With R. H. Zander P. M. Eckel 431285 April 5, 1985 (BUF).

 

CAN. gorge of Niagara River, Niagara Glen, near path, floor of woods near river, moist rock surface, R. Zander & P. Eckel, Dec. 4, 2004 (MO).

USA: just outside spray area of Horseshoe Falls, boulder, thin soil, midslope, with Tortella fragilis, Hymenostylium recurvirostrum, Trichostomum crispulum, Zander 3484c Oct. 28, 1979 (BUF), Zander 3480, Oct. 28, 1970 (BUF); spray area of Horseshoe Falls, near river, thin soil in crevice of boulder, Zander 3469, Oct. 28, 1970 (BUF).

Hypnum abietinum Hedw. = Thuidium abietinum (Hedw.) BSG

Hypnum aduncum Hedw. = Drepanocladus aduncus (Hedw.) Warnst.

Hypnum alleghaniense C.Muell. = Thamnobryum alleghaniense (C. Muell.) Nieuwl. 

 

Hypnum brevirostre Ehrh. A Clinton specimen from Devil's Hole (herb no. 41058).

 

Hypnum confervoides Schwaeg. Schwaegr.

 

USA: On rocks, with Radula complanata, second sister Goat island (1873  Clinton, Coll. herb. no. 41072 (BUF).

 

Hypnum cupressiforme Hedw.  NY CHECK

 

USA: Artpark, Eckel 451486 (BUF), (Eckel, l986b).

 

Hypnum curvifolium Hedw. NY CHECK

 

USA: Suspension Bridge (now City of Niagara Falls by the Lower Arch Bridge) archived letter BUF from Charles Peck to George Clinton (collector, from Clinton's  trip May 11, 1867 noted in Clinton's journal)

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring); P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

USA: Erie Co. Grand Island. At delta of creek flowing into Niagara R,

south tip of Grand Island. Ann Glowny 1096 3-28-67(BUF).

 

Hypnum imponens Hedw. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: Navy Island in the Niagara River just S of Niagara Falls, NW end, open gallery, spring-wet Quercus, Ulmus americana, Acer rubrum woods.P. M. Eckel s.n. July 2, 1998(BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Fallen log; embankment; with Dicranum fuscescens.  Pseudoparaphyllia laciniate. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF); Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. W side; fallen log; with Ptilidium pulcherrimum, Dicranum flagellare. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

 Note this species has costa short and double, and that its stem cortical cells are little or not inflated but especially the few thicker‑walled orange cells at the base of the leaf subtended by a few enlarged thin‑walled hyaline cells (cf. Drepanocladus, which has a costa).

 

Hypnum lindbergii Mitt. (New to NY CHECK)

 

USA: Devil's Hole, upper slopes, Eckel 331585 (BUF, NYS), Eckel, l986a; wet calcareous flat at base of gorge in shrubby river margin, P. M. Eckel Sept. 22, 2004 (BUF); Devil's Hole by the Power Authority, Niagara River Gorge. Upper slopes, calcareous, with Dryopteris marginalis, on rock talus, with

With R. Zander. Dicranella varia. P. M. Eckel 331585 March 16, 1985

(BUF).

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. On punky log in late snow melt - boulder field. Note there are few rotten log substrates in the flats. With R. Zander P. M. Eckel 531285 April 5, 1985 (BUF).

 

USA: Second of the Three Sisters Islands, east end, on the Niagara River just S of Goat Island. Northern margins by water. P.M. Eckel 881111 Aug. 8, 1988 (BUF); Second Sister, Eckel 86121701 (BUF, Bailey)

 

USA: Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring), P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

CAN: Dufferin Islands, marshy area on log, Oct. 8, 1989, Eckel  9303175 (BUF).

USA: Second Sister Island, west end, 2 Nov. 1988, Buck 16379 (BUF, NY).

USA: Second Sister Island, east end, thin soil over dolomite flats, open with Salix, Cornus, Betula, Lythrum, Carices, Eckel 86121701, Sept. 12, 1986 (BUF); 2 Nov. 1988, Buck 16404 (BUF, NY).

CAN: Navy Island in the Niagara River just S of Niagara Falls, northwest and northern river margins; "slump meadow" wet red clay eroding embankment. P. M. Eckel s.n. July 15, 1998 (BUF); Navy Island in the Niagara River: 43*03'N,79*01'W, just south of Niagara Falls. Dominant on soil, island west side; Beech-Maple, Carya-Quercus forest; Atrichum undulatum. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF); Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Dominant on logs, interior of island, with Dicranum flagellare; Red Maple forest, wet. P. M. Eckel with R. Zander s.n. Sept. 14, 1998(BUF).

 

Hypnum pallescens (Hedw.) P. Beauv.

USA: Second Sister Island, west end, on trunk of Salix, 2 Nov. 1988, Buck 16388 (BUF, NY).

CAN: Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. With Hedwigia ciliata on granite glacial erratic; Fraxinus-Carya woods P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Hypnum pygmaeum (BSG) Sull. & Lesq. = Thuidium pygmaeum BSG [note also Hypnum pygmaeum Vill. ex Brid. = Leskeella nervosa (Brid.) Loesk.

 

Hypnum rivulare Brid.(BSG) Bruch =  Brachythecium rivulare  BSG

 

Hypnum rugosum Ehrh. ex Hedw. = Rhytidium rugosum (Hedw.) Kindb.

 

 Hypnum sprucei (Bruch) Spruc. = Amblystegium jungermannioides (Brid.) Giac. fid. Giac.

 

Hypnum subcompactum,  a doubtful name, not in Index Muscorum

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: "... on rocks and stones below Queenston Heights, Ont. (Macoun.)," Macoun, 1902, p. 312.  CHECK CITATION

 

Hypnum schreberi Willd.

 

[CAN?]: City Niagara Falls. Wood below the Whirlpool. Ex Coll. G. W. Clinton, Buffalo, N.Y. (BUF).

 

A specimen from "wood below the Whirlpool" which Zander has suggested is the wooded moss-rock house area associated with Whirlpool (DeVeaux) steps.

 

Hypnum sullivantii R. Spruce

 

A Clinton specimen from Devil's Hole with fruit (herb no. 38547).

 

Isopterygium elegans (Brid.) Lindb. NY CHECK

 

Kindbergia praelonga (Hedw.) Ochyra

 

As Stokesiella praelonga (Hedw.) Robins.

 

Found by Macoun in 1891 at Foster's Flats, Welland Co. (CANM 183141). Note other records in Cook, 1985. (Crum & Anderson, Ireland etc.)

 

(As Stokesiella praelonga (Hedw.) Robins. in Crum & Anderson, l981.

 

See Crum & Anderson 1981, p. 1077: Canada, Foster's Flats.

 

Leptobryum pyriforme (Hedw.) Wils.

 

CAN: As "Leptobigum pyriforma," Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: Luna Island, Niagara Falls [cf. handwriting E. G. Britton, A. A. A. S. meeting] Aug. 21, 1886 (NY, in packet of Funaria hygrometrica). 

 

USA: Buckhorn Island State Park.

 

USA: New York, Erie Co., Grand Island Twp., NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring). P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF), with Pohlia nutans.

 

CAN: Leptobryum vulgare, Queen Victoria Niagara Falls Park.  Cameron 1895.. Doubtful name. Possibly Leucobryum vulgare (= L. glaucum) was meant.

 

Leptodictium humile (P.Beauv.) Ochyra

 

CAN: Ontario, Regnl. Mun.   Niagara, town  of Niagara‑on‑the‑Lake, W side Niagara River on S boundary Fort George. Spring‑inundated soil sub Quercus alba, Quercus rubra forest,  with Brachythecium campestre and Physcomitrium pyriforme. P. M. Eckel, Oct. 4, 1997 BUF.

 

Leptodictium riparium (Hedw.) Warnst.

(As Amblystegium riparium). USA: Second Sister Island, west end, tree roots, wet in high water, Oct. 29, 1988 (BUF).

(As A. riparium) Goat Island in the Niagara River, southeast end, wet soil by riverside. Open, in mats. P. M. Eckel June 3, 1987 (BUF); Dolomite ballast, open eastern end of Goat Island, south side. Barbula unguiculata,Bryum lisae var cuspidatum P.M. Eckel June 4, 1988 (BUF).

 

Leptodictyum trichopodium var. kochii (BSG) Broth.                     

 

Amblystegium kochii, Bruch & Schimp.

 

Leptodictyum trichopodium var. curvipes (BSG) Broth. is reported as excluded from Canada, Ireland et al. mosses of Canada. Macoun (1902) reported the var. curvipes at Niagara Falls, but gave Amblystegium kochii in its synonymy. What was probably found at Niagara was the var. kochii

 

As Amblystegium curvipes BSG "On old logs, Foster's Flats, Niagara Falls, Ont., May 13th, 1901 ... (Macoun.)," Macoun, 1902, p. 302. 

 

Lescuraea patens (Lindb.) H. Arnell & C. Jens.

 

Pseudoleskea atrovirens (Brid.) BSG

 

As Pseudoleskea atrovirens, "On rocks at Niagara Falls (Macoun.)," Macoun & Kindberg, 1892, p. 180. 

 

As Hypnum atrovirens Dicks. and Hypnum filamentosum, Dicks. l.c.

 

Pseudoleskea atrovirens Bruch & Schimp.

 

"On rocks; Niagara Falls and Lake Superior (Macoun)..." (Lesquereux & James, 1884, p. 319), probably in Canada.

 

Leskea gracilescens Hedw.

 

USA: Erie Co. Grand Island. Marshy oak woods on Love Rd., Niagara R.

Tree base in swamp. Ann Glowny 1363 4-20-67 (BUF).

 

Leskea nervosa (Brid.) Myr. = Leskeella nervosa (Brid.) Loeske 

 

Leskea polycarpa Hedw. var. polycarpa NY CHECK

 

Hypnum medium, Drumm. Musc. Bor.‑Am. No. 223.

 

Leskea subobtusifolia C. Muell. & Kindb. in Mac. & Kindb. 1892.

 

USA: RR Bridge, Upper path Eckel 33895 (BUF), Eckel l986.

 

USA: Erie Co., north end of Grand Island, Buckhorn Island State Park, SW portion of Park, west of I 190, near path along Niagara River from West River Parkway to Burn Ship Creek, on bark of thick Populus deltoides branch at face height, near two excavated areas in Typha latifolia marsh. P. M. Eckel, Oct. 1, 1997, with R.  H. Zander, with Ceratodon purpureus, Orthotrichum obtusifolium (BUF). 

 

New York: Buckhorn Island, east end, east shore of Burnt Ship creek, on Ulmus americana, with Platygyrium repens, Orthotrichum ohioense, O. obtusifolium, Eckel Oct. 21, 1997 (BUF). 

USA: First Sister Island, western half, 2 Nov. 1988, Buck 16419 (BUF, NY).

USA: Whirlpool St. by the RR Bridge. Upper path along rim of gorge along dolomite caprock. With Orthotrichum anomalum, Bryum capillare. With R. Zander. P. M. Eckel 33895 March 9, 1985 (BUF).

 

USA: Erie Co. Grand Island. Marshy oak woods, Bflo. Ornithological Soc.

woods, Niagara River. Ann Glowny 1358 4-20-67 (BUF).

 

CAN:  Navy Island in the Niagara River, northeast side, shore, fallen tree on beach, bark on trunk; with Entodon seductrix, Ceratodon purpureus. P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF).

 

Leskea polycarpa Ehrh. var. paludosa (Hedw.) Schimp.

 

"On trees, Niagara Falls, Ont., June 15th, 1884 ..." Macoun, 1902, p. 168.  "The above were referred to L. polycarpa in Part VI, p. 168," Macoun, 1902, p. 272. In Part VI (1892) L. polycarpa was

 

"… abundant on trees, Foster's Flats, below Niagara Falls, (Macoun.)."

 

Note that there are only literature reports for this variety as given by Ireland et al. (1980). 

 

Leskeella nervosa (Brid.) Loeske NY CHECK

 

note Hypnum pygmaeum Vill. ex Brid. = Leskeella nervosa (Brid.) Loesk. Hypnum pygmaeum (BSG) Sull. & Lesq. is Thuidium pygmaeum BSG

 

= Leskea nervosa (Brid.) Myr.

 

USA: (As Leskea nervosa)  "Rocks. Niagara Falls, G. W. Clinton. … The specimens are sterile, but believed to belong to this species." (Peck 1866).

 

(As Leskea nervosa), Niagara Falls., Day 1883.

 

Leucobryum glaucum (Hedw.) Aongstr. ex Fr.

 

Leucobryum vulgare Hampe

 

(As Leucobryum vulgare) "... woods, Queenston Heights, Ont. (Macoun.)," Macoun, 1902, p. 204‑05. 

 

Mnium affine Bland ex Funck

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

Mnium affine Bland ex Funck = Plagiomnium affine (Funck.) Kop. "Excluded from Canada. Most collections thus named are either P. ciliare or P. ellipticum" (Ireland, et al., 1980.)

 

Mnium ciliare (C. Muell.) Schimp. NY CHECK

 

USA: (as Mnium affine var. ciliare) Goat Island, south side, flats just west of Three Sisters (Eckel 1990b); Dolomite flats just west of the Three Sisters Islands. With Brachythecium salebrosum. det. R. Zander P. M. Eckel July 5, 1988 (BUF).

 

CAN: (as Mnium affine var. ciliare) Navy Island in the Niagara River: 43*03'N,

79*01'W, just south of Niagara Falls, with Atrichum undulatum var. oerstedianum; hummock Beeches dominant. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF).

 

Mnium cuspidatum Hedw. NY CHECK

 

USA: Suspension Bridge (now City of Niagara Falls by the Lower Arch Bridge)  archived letter BUF from Charles Peck to George Clinton (collector, from Clinton's  trip May 11, 1867 noted in Clinton's journal)

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: Near the top of Whirlpool Steps, N of the Whirlpool, Ongiara Gorge.

 w. R. Zander. P. M. Eckel 880924 Sept. 5, 1988 (BUF).

 

USA: Devil's Hole on the Niagara River. Upper slopes, calcareous bedrock. Soil in trash-midden. Plants synoicous. With R. Zander. P. M. Eckel 132185 March 16, 1985(BUF). 

 

USA: City of Niagara Falls, Whirlpool State Park, in solo sylvatico sterile calcareo, in luco Querci rubri, umbroso, P. M. Eckel, Oct. 4, 1997 (BUF), with Amblystegium varium.

 

CAN: Niagara Glen (Foster's Flats) Just N Of The Whirlpool, Base Of Dolomite Boulder, N-Facing Aspect, With Lophocolea Heterophylla, Myurella Sibirica. P. M. Eckel 521186 April 11, L985 (BUF); Niagara Glen, just N of Whirlpool. Shaded soil on limestone boulder, base of seedling tree, Acer - Tsuga canadensis wood. With R.H.Zander. P.M.Eckel NF6 Feb. 19, 1983 (BUF).

 

CAN: Navy Island in the Niagara River, northeast side, dry clay embankment, with Brachythecium rutabulum. P. M. Eckel with R. Zander s.n. July 15, 1998  (BUF); Navy Island in the Niagara River, northeast side, dry fringe meadow; soil, characteristic of vegetation of meadow between crest and shrubby forest border. P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Soil; top of shore embankment; shady; dominant moss. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

USA: Goat Island, south side, flats just west of Three Sisters, wet hummock, roots of Cornus (Eckel 1990b); Goat Island on the Niagara River. SW end, shoreline, flats east of the Horseshoe Falls, wet hummock, roots of Cornus stolonifera. P. M. Eckel 880716 June 3, 1987 (BUF).

 

USA: (as Plagiomnium cuspidatum.) Second Sister Island, west end, 2 Nov. 1988, Buck 16390 (BUF, NY); (as Plagiomnium cuspidatum); Second Sister Island, east end, 2 Nov. 1988, Buck 16401 (BUF, NY).

USA: Third Sister Island, west end, in vug of dolomite boulder. Eckel & Eckel 8703252, March 3, 1987 (BUF).

Mnium marginatum (With.) Brid ex P.‑Beauv. NY CHECK New!

 

In Ontario, it appears that the Whirlpool woods area, perhaps in Bowman's Ravine, is a type locality:

 

"Mnium niagarae Kindbarg in Macoun & Kindbarg, Catalogue of Canadian Plants 6: 141. 1892. Isotypes. Canada: Ontario, Niagara Falls, Whirlpool, 6 June 1891, Canadian Musci [no.] 581 (CANM 198223); Ontario, Niagara, Whirlpool, 6 June 1891 (CANM 19822). = M. marginatum (With.) P. Beauv. (Ireland  & Ley 1981).

 

CAN: (As Mnium niagarae Kindb.) "On earth in woods, Whirlpool, Niagara Falls, June 6, 1891" (Crum 1981).

 

CAN: (As Mnium niagarae Kindb. in Macoun), Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: (As Mnium niagarae Kindb. ex Macoun & Kindb.), p. 141, Cat. Canad. Pl. vol. 6. 1892.

 

(As Mnium Niagarae, Kindb. in Mac. & Kindb.) Canadian Musci, No. 581 "Intermediate between M. spinulosum and M. serratum, but readily distinguished from both by the wide‑mouthed capsule." "On earth under trees near the water in the woods bordering the Whirlpool, Niagara Falls, Ont., June 6th, 1891. (Macoun.)," Macoun & Kindberg, 1892, pp. 141‑42. 

 

As "M. Niagarae Kindb. in Macoun & Kindb., Cat. Canad. Pl. 6: 141. 1892. Holotype: Ontario. Niagara Falls, on earth under trees near the water in the woods bordering the Whirlpool, 6 VI. 1891 Macoun (+Macoun, Canad. Musci exs. 581) (S‑PA, isotypes in CANM and H‑BR). ... It often has been collected together with M. thomsonii on calcareous cliffs." p. 168. Koponen (1974).

 

CAN: Niagara Glen, just N of Whirlpool. semi shaded vertical face of limestone boulder. Thin soil in pockets. Synoicous. With R.H.Zander. P.M.Eckel NF13 Feb. 19, 1983 (BUF). 

 

USA: Zander, Ongiara, lower slopes

 

USA: Devil's Hole Near The Top On Dolomite Or Lime-Stone Talus, Rather Arid Situation, With Dryopteris Marginalis, Fissidens Taxifolius, Ceratodon purpureus. P. Eckel 821186 March 16, L985 (BUF).

 

Mnium medium BSG NY CHECK

 

Mnium orthorrhynchum Bruch & Schimp.

 

"Niagara Falls (Clinton)..." (Lesquereux & James, 1884, p. 246).

 

CAN: "Crevices of limestone rocks, Niagara Falls, 1882, …" [Macoun specimen] (Crum 1891).

 

Mnium punctatum Hedw. var. punctatum NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. S end, decorticated log, Oak-Hickory; with Nowellia curvifolia. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Mnium rostratum Schrad. NY CHECK

 

As Mnium rostratum Schrad. Devil's Hole, Niagara River. Day 1883.

 

Mnium spinulosum Bruch & Schimp.

 

"... in woods, Niagara River at Foster's Flats, Ont., May 13th, 1901. (Macoun.)," Macoun, 1902, p. 258.

 

Mnium thomsonii Schimp. NY CHECK

 

CAN: Eckel, Foster's Flats, halfway down.

 

USA: Whirlpool steps 1/4 mile N of Whirlpool Park, half way down gorge of Niagara River, Thuja clump, dolomite talus, west facing, N aspect,  with Campylium chrysophyllum, Tortella tortuosa. May 6, 1987, Eckel No. 9302092 c (BUF) Note this sp. seems absent from the deciduous zone in Ontario boulder field. Koponen says M. thomsonii "has a very narrow ecological amplitude; it grows exclusively on limestone or calcareous cliffs, or on calcareous soil in arctic conditions" p.  166. [Koponen, T.  Lindbergia 2:160‑184].

USA: First Sister Island, western half, Eckel 880720, July 8, 1987 (BUF); 1 Nov. 1988, Buck 16338 (NY).

Myurella sibirica (C. Muell.) Reim. NY CHECK

 

USA: Zander, Ongiara, lower,  Zander 3517 BUF, NYS  (Eckel, l986b).

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Far back in totally sheltered niched of dolomite boulders, with Mnium thomsonii. With R. H. Zander    Late snow melt. P. M. Eckel  13885 April 5, 1985(BUF CANM) Niagara Glen (Foster's Flats) Just N Of The Whirlpool, Base Of Dolomite Boulder, N-Facing Aspect, With Lophocolea Heterophylla, Mnium Cuspicatum. P. M. Eckel 421186 April 11, L985 (BUF). 

 

Neckera pennata Hedw.

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

 Orthotrichum anomalum Hedw.

 

One specimen of Drummond's exsiccat No. 152 at NY contained Orthotrichum anomalum, rather than the O. cupulatum indicated on the label: " Rocks near the Whirlpool, below the Falls of Niagara."

 

USA: "Limestone rocks, Niagara Falls, Lesqx. On Trees in the same locality, G. W. Clinton." (Peck 1866).

 

"Rare. ..Niagara Falls," Day 1883.

 

"On limestone rocks; Niagara Falls ..." (Lesquereux & James, 1884, pp.164‑165).

 

Orthotrichum anomalum Hedw.  "... on perpendicular rock, Foster's Flats, Niagara River, 1901.  (Macoun.)," Macoun, 1902, p. 229. 

 

USA: Niagara Falls, N.Y. Charles Mohr, Coll. July 1, 1866, herbarium of Charles Mohr, presented in 1901, "ad saxas calcareas," c. fr. (US).

 

USA: No. 119 S. & L. M. N. Amer. ed. I. "Niagara" Agassiz (NY).  label hand written.

 

USA: No. 177 Musci Boreali‑Americani, W. S. Sullivant & L. Lesquereux, ed 2. Ad rupes Niagara (Agassiz) (NY). label hand written. Also, with printed label.

 

USA: New York, Niagara Falls (2), also Whitehall (1) and Albany (3), "1 & 3 C. H. Peck. 2. G. W. Clinton" (NYS)

 

USA: New York, Niagara Falls and vicinity, G. W. Clinton (NYS).

 

USA: Musci Boreali‑Americani, W. S. Sullivant & L. Lesquereux, MDCCCLVI No. 119, ed. 1 Hab. ad rupes Niagarae cataractae (Agassiz) (NY). 

 

USA: New York. Niagara Falls. specimen says H. Gillman, Coll. label, however, of Charles H. Peck showing Clinton as the collector is included in the specimen (NY). 

 

CAN: On rocks. Niagara River. Mosses from Ontario, Canada, J. Macoun, May 13, 1901 (NY).

 

[USA?] Hab. Niagara, 1873, Francis Wolle c.fr. (NY).

 

[USA?] Hab. Niagara, trees above Falls. Herb. Eugene A. Rau, Bethlehem, Pa. (NY).

 

USA: At Whirlpool St by the RR bridge under R. Moses Pkwy, path  along the dolomite caprock. Vertical surface of a dolomite rock as part of talus slope, mesic, rather shady. PM Eckel & RH Zander 331685 Mar 9 1985 (BUF).

 

CAN: Niagara Glen, just N of Whirlpool. Thin soil over calcareous boulder - shaded in summer. With R.H.Zander P.M.Eckel NF3 Feb. 19, 1983 (BUF).

 

CAN: Whirlpool Ravine, base of shaded dolomite, June 9, 1990, c.fr., deoperculated, Eckel 9302096 (BUF).

 

USA: Devil's Hole, upper slopes, Eckel 526386.

 

USA: Goat Island, south shore, on boulders toward the Three Sisters (Eckel 1990b); Goat Island on the Niagara River, wooded slope just N of first 3-Sisters Islds Bridge, on boulder.  c.fr. P. M. Eckel 881005 Sept. 30, 1988(BUF).

USA: First Sister Island, western half, Eckel 880721, July 8, 1987 (BUF).

USA: Green Isl. on the Niagara R, just N of Goat Isl.  Old dolomite ballast on S side, wet to moist river margin. With Grimmia sp. P. M. Eckel, Marguerite P Eckel. 9004154 April 15, 1990 (BUF).

 

Orthotrichum cupulatum Brid. NY CHECK  Doubtful reports.

 

Perhaps this literature report refers to Day's citation below. Vitt suggests that eastern reports for O. cupulatum are O. strangulatum, as the former species occurs in the west. 

 

As Orthotrichum cupulatum Hoff.."Niagara Falls, Drummond," Sullivant, Charles Peck in Day 1883.

 

As Orthotrichum nudum Dicks. var. Rudolphianum (Lehm.) Husn. "Rocks near the Whirlpool, below Niagara Falls. (Drummond.)," Macoun & Kindberg, 1892, p. 87. 

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Seepage just below the dolomite caprock of the Niagara gorge. With R. H. Zander P. M. Eckel 321185 April 5, 1985 (BUF).

 

Orthotrichum obtusifolium Brid.

 

USA: Erie Co., north end of Grand Island, Buckhorn Island State Park, SW portion of Park, west of I 190, near path along Niagara River from West River Parkway to Burn Ship Creek, on bark of thick Populus deltoides branch at face height, near two excavated areas in Typha latifolia marsh. P. M. Eckel, Oct. 1, 1997, with R. H. Zander, with Ceratodon purpureus.

 

Orthotrichum obtusifolium Brid.

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

Orthotrichum ohioense Sull. & Lesq. ex Aust.

 

2005 Rare Plant Bryophyte Status List SH

 

CAN: base of wooded embankment overlooking the Horseshoe Falls, in spray on mature Salix, Eckel & Zander, c.fr. May 27, 1991, No. 9302251 (BUF).

 

New York: Buckhorn Island, east end, east shore of Burnt Ship creek, on Ulmus americana, with Platygyrium repens, Leskea polycarpa, O. obtusifolium, Eckel Oct. 21, 1997 (BUF). 

 

This species is distinct from O. pusillum, especially when the latter is old and ribbed, by the reflexed teeth when dry, and the smaller leaf cell walls that are incrassate and irregularly thickened. The cells of O. pusillum are not incrassate and have even walls.  The calyptra of O. ohioense is hairy, that of O. pusillum smooth. The brood bodies in material from Buckhorn Island are from a few stems of O. obtusifolium in the population. Both specimens were on tree trunks in the open (without other canopy).  The species grows on the bark of deciduous trees including Acer, Fraxinus, Carpinus and Cornus, all of which grow at Niagara. Since populations do not usually grow in the open, the trees of the Niagara collections may be assumed to have once been on the forest edge but have been exposed by park personnel. 

 

Orthotrichum speciosum Nees ex Sturm.

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

Orthotrichum strangulatum P.‑Beauv.

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

As Orthotrichum Porteri, Sull. & Lesq. in Aust., Bull. Torr. Bot. Club, Vol.  VI, 34 "On dry limestone rock, by the stairs leading down the cliff to Foster's Flats, Niagara Falls, Ont., May 16th, 1901. (Macoun.)," Macoun, 1902, p. 230. 

 

"Some of the material distributed as Drummond's Musci Americani (Rocky Mts.) no. 152, as O. cupulatum, is indeed that species of western distribution; some is O. strangulatum instead. It is doubtful if O. cupulatum was actually collected at Niagara Falls, as the label indicates, but O. strangulatum could have been" (Crum & Anderson 1981, p. 707).

 

Phascum cuspidatum Hedw. NY CHECK

 

=Tortula atherodes Zander

 

"On earth on the wood roads around the base of Queenston Heights, Ont.; in fine fruit, May 8th, 1901. Found only on hard baked earth. (Macoun.)," Macoun, 1902, p. 185. 

 

Ontario: Drummond's exsiccat Musci Boreali‑Americani, number 6. 1828, as var. elatum Hooker et Tayl. Musc. Brit. ed 2, p. 9] "Hab. Sandy field near the Falls of Niagara" [NY, no specimen accompanied the label]. Grout, in his treatment of Phascum, considered Drummond's No. 6 to be var. Americanum Ren. & Card. The typical variety was considered by him to occur in the western United Stated (Washington), the var. americanum to be "widely distributed in southern Canada" &c.

 

CAN: Ontario, On earth, roadside, below Queenston Heights, 8/5/01, J. M. Macoun, No. 200, May 8, 1901 (NYS).

 

USA: Zander, Eddy Basin, upper slopes.

 

CAN: Niagara-on-the-Lake. across the Niagara Pky (west) and S of Fort George. Paradise Grove; hard clay, disturbed dirt road, garden midden. P. M. Eckel with R. H. Zander s.n. May 25, 1998 (BUF), with Physcomitrium pyriforme, Weissia sp.; Niagara-on-the-Lake. Soil, between grasses of lawn. Richard H. Zander 4221 June 22, 1974 (BUF).

 

CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Dicranella varia, Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Barbula unguiculata, May 27, 1991, Eckel 9302250 D (BUF). 

 

CAN: Dufferin Islands near the Falls, western area, bare soil in lawn. R. Zander & P. Eckel 8612281 Oct. 30, 1986 (BUF).

 

USA: Goat Island on the Niagara River. Soil at base of tree by the restaurant beside the driveway. With Barbula unguiculata and Pottia truncata var. truncata. P. M. Eckel 881030 Oct. 29, 1988 BUF); Goat Island on the Niagara River. North side in island of roads "by maintenance building" base of Acer saccharum. P.M. Eckel 881115 Nov. 10, 1988 (BUF).

 

USA: Green Island on the Niagara River, just N of Goat Island. Old dolomite ballast on south side, wet to moist river margin. PM Eckel, Marguerite P. Eckel 9004152 April 15 1990  (BUF).

 

USA; New York: Erie Co. Grand Island. Marshy soil by road, southern end of Grand Island, Niagara River. Ann Glowny 1156 Mar. 28, 1968 (BUF).

 

Philonotis fontana (Hedw.) Brid.NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: "... in damp woods by Drummond's Island, Niagara Falls, Ont., May, 1900. (Macoun.)," Macoun, 1902, p. 240.

 

[USA?] Niagara Falls. as var. falcata [Bartamia fontana var. falcata. Rev. F. Wolle, 1873, herb. Coe Finch Austin (NY)

 

CAN: Niagara River Gorge, near Horseshoe Falls, near bottom of slope, spray zone, on thin soil on rocks with grasses det. P. M. Eckel  1995 R. H. Zander 3696 Sept. 7, 1971 (BUF).

 

Philonotis marchica (Hedw.) Brid.

 

USA: Gorge of Niagara River, near the American Falls by Prospect Point, near river, thin soil, crevices of limestone boulder. det. P. M. Eckel  1995 R. H. Zander 3741c Sept. 30, 1971(BUF).

 

USA: Goat Island, Eckel 216686 (BUF, NYS), (Eckel, l986b).

USA: Goat Island, Terrapin Point, 2 Nov. 1988, Buck, 16439 (BUF, NY).

USA: Second Sister Island, east end,  2 Nov. 1988, Buck 16394 (NY).

Philonotis muhlenbergii (Schwaegr.) Brid NY CHECK

 

2005 Rare Plant Bryophyte Status List SH

 

= Bartramia muhlenbergii Schwaegr.

 

USA: (As Bartramia muhlenbergii (Schwaegr.) Brid.) "Niagara Falls, G. W. Clinton." (Peck 1866),

 

As Bartramia muhlenbergii Schwaegr. Niagara Falls, Day 1883.

 

According to Crum and Anderson 1981: "Cardot (Bull. Herb. Boiss. 7:307, 1899) was unable to distinguish the type from Philonotis marchica.. The problems in species definition are so difficult in Philonotis that we have avoided taking a definite stand.... William Zales, in his unpublished dissertation on Philonitis in North America (of which we have seen the key to species), apparently recognized P. marchica but not P. muhlenbergii. Other authors have expressed doubt that the two species can be differentiated."

 

One specimen I originally identified as P. muhlenbergiana, but upon examination of other stems I find that lower leaves are subpercurrent, and in this collection, at least some of the leaves had percurrent costae, hence the determination P. marchica (Eckel 216686) above.

 

USA: Goat Island, south side, flats just west of Three Sisters, wet hummock, roots of Cornus (Eckel 1990b); Goat Island on the Niagara River. SW end, shoreline, flats east of the Horseshoe Falls, wet hummock, roots of Cornus stolonifera'; lvs percurrent (vs. P. marchica). M. Eckel 880718 June 3, 1987 (BUF).

USA: Second Sister Island, west end, tree roots, wet in high water, Oct. 29, 1988 (BUF).

Physcomitrium pyriforme (Hedw.) Hampe

 

As Physcomitrium turbinatum C. M. ex Lesq. & James, Man. Mosses No. Amer., p. 198, 1884, non  (Mx.) E. G. Britt. 1894.

 

As Physcomitrium turbinatum C. Muell. "Very abundant on earth below Queenston Heights and within the spray of Niagara Falls, Ont., above the Power House, and on earth in the flower border close to the falls, May 16th, 1901. (Macoun.)," Macoun, 1902, p.  238. 

 

CAN: Ontario, Regnl. Mun. Niagara, town of Niagara‑on‑the‑Lake, W side Niagara River on S boundary Fort George. Spring‑inundated soil sub Quercus alba, Quercus rubra old forest.  P. M. Eckel, Oct. 4, 1997 (BUF), c.fr., with Brachythecium campestre, fruit young, & Leptodictium humile c.fr.; also dirt road, with Phascum cuspidatum, Weissia sp. Eckel May 25, 1998 (BUF).

 

CAN: base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Barbula unguiculata, May 27, 1991, Eckel No. 9302250C May 27, 1991 (BUF). 

 

USA: Bottom of slope near Sewage Treatment Plant, Eckel 125586 (BUF), (Eckel, l986b).

 

USA: Upper Great Gorge of the Niagara River. Just S of the sewage treatment plant on disturbed slope, with grasses, cinder substrata. With Weissia controversa, Astomum muhlenberg. P. M. Eckel 125586 May 24, l986 (BUF).

 

A "winter annual and springtime weed" (Crum & Anderson, p. 464), "It is a weed and could well have been introduced from Europe, and it shows a Crataegus‑like evolutionary burgeoning in North America," its existence in the Niagara parklands, especially so early (1901), may be an indication of tourist‑effect, much like the report of Bryum rubens reported above. 

 

Plagiopus oederiana (Sw.) Limpr. NY CHECK

 

Bartramia oederi Brid.

 

Bartramia oederiana Turn.

 

USA: (As Bartramia oederi Swartz.) "Devil's Hole, G. W. Clinton." (Peck 1866).

 

USA: (As Bartramia oerderi Swartz.) Devil's Hole, Niagara River, Day 1883.

 

CAN: (As  Bartramia oerderiana) Queen Victoria Niagara Falls Park .Cameron 1895.

 

USA: Zander, Ongiara, lower slopes Zander 3521c (BUF, NYS), (Eckel, l986b).

 

Plagiothecium cavifolium (Brid.) Iwats.

USA: Second Sister Island, west end, 2 Nov. 1988, Buck 16370 (NY), 16376 (BUF, NY).

Plagiothecium denticulatum (Hedw.) BSG

 

CAN: Queen Victoria Niagara Falls Park .Cameron 1895.

 

CAN: "... also at Niagara Falls ... 1901 (Macoun.)," Macoun, 1902, p. 299.

 

USA: Goat Island, east end ballast on island margins (Eckel 1990b); Goat Island on the Niagara River. Dolomite ballast, southeast end, just east of the Three Sisters Islands. det. R. H. Zander P. M. Eckel May 22, 1988 (BUF).

USA: First Sister Island, western half, 2 Nov. 1988, Buck 16424 (BUF, NY).

Plagiothecium latebricola BSG

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring); P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF); cortical cells large & thin, lvs in decurrencies with propagula. The plant was found in the Quercus alba, Q. rubra, Q. palustris and Acer rubrum swampy woods of Buckhorn Island State Park on a punky log in soil subject to flooding. Crum and Anderson (1981) indicate that the species is rare in North America, although Ketchledge (1980) records numerous stations in New York State. It is separated from Isopterygium by the thin‑walled cortical cells and the enlarged and clear cells in the leaf angles being distinctly decurrent. The stems and branch cortical cells of Isopterygium are small and thick‑walled, the leaves have no decurrencies even though the filiform propagulae seem similar to those of species in Plagiothecium.

 

CAN: Gorge of Niagara River, near the American Falls by Prospect Point, near river, thin soil, crevices of limestone boulder.  R. H. Zander 3741c Sept. 30, 1971 (BUF).

 

Platydictya confervoides (Brid.) Crum

 

USA: Niagara Co., Whirlpool woods, G. W. Clinton (NYS) [glued on card: note the identification of Whirlpool woods with a New York station].

 

CAN: Niagara Glen, just N of Whirlpool. Thin soil over limestone boulder, shade. With Anomodon rostratus, Encalypta procera, Tortella tortuosa. With R.H.Zander. P.M.Eckel NF1 Feb. 19, 1983(BUF); Niagara Glen, just N of Whirlpool. Thin soil over limestone boulder, cool, bottomland Acer - Tsuga canadensis wood. With R.H.Zander P.M.Eckel NF8 Feb. 19, 1983 (BUF); Gorge of Niagara River, Niagara Glen, half

way up slope from river, Pot Hole, Path, boulder, thin soil. R. H. Zander 3730 Sept. 29, 1971 (BUF).

 

USA: Whirlpool steps, base of talus slope, with Didymodon rigidulus var. rigidulus, Campylium chrysophyllum. Ongiara Gorge. w. R. Zander. P. M. Eckel 880929 Sept. 5, 1988 (BUF).

 

Platydictya jungermannioides (Brid.) Crum. NY CHECK

 

As Hypnum sprucii Bruch. on the ground. James. Goat island Falls Niagara NY  July 1850 herb. no. 41186.

 

Platydictya minutissima (Sull. & Lesqu. ex Sull.) Crum NY CHECK

 

2005 Rare Plant Bryophyte Status List S3? [sic]

 

Platydictya subtile (Brid.) Crum NY CHECK

 

As Hypnum subtile Hoffm.  Whirlpool woods. Day 1883. 

 

[CAN?] City Niagara Falls. Whirlpool Woods. Ex Coll. G. W. Clinton, Buffalo, N. Y. June 10 (BUF); Clinton specimen for Whirlpool woods, June 14 herb. no. 41200 [1980-1870].

 

Platygyrium repens (Brid.) BSG NY CHECK

 

USA: Artpark, Eckel & Eckel 251486 (BUF).

 

USA: Whirlpool Creek [Devil's Hole, creek is called "Bloody Run"] G. W. Clinton [glued on card] [NYS]

 

New York: Buckhorn Island, east end, east shore of Burnt Ship creek, on Ulmus americana, with Leskea polycarpa, Orthotrichum ohioense, O. obtusifolium, Eckel Oct. 21, 1997 (BUF). 

USA: Second Sister Island, west end, on rotten log, 2 Nov. 1988, Buck, 16369 (BUF, NY).

CAN: Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Bark of Fraxinus pensylvanica. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

Pleuridium subulatum (Hedw.) Rabenh.

 

CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Dicranella varia, Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Barbula unguiculata, May 27, 1991, Eckel No. 9302250 E (BUF). 

 

Pohlia annotina (Hedw.) Lindb.

 

CAN: As Webera annotina (Hedw.) Bruch in Schwaegr., Queen Victoria Niagara Falls Park .Cameron 1895.    

 

Pohlia atropurpurea (Wahl.) H. Lindb.

 

This species is on the 2005 Rare Plant Bryophyte Status List SH

 

Bryum atropurpureum Wahlenb. in Fuernr.

 

CAN: As Bryum atropurium (= atropurpureum Wahl. ex Web. & Mohr), Queen Victoria Niagara Falls Park .Cameron 1895.

 

CAN: As Bryum atropurpureum Wahlenb. in Fuernr.  "On earth at Foster's Flats below Niagara Falls, Ont. (Macoun.)," Macoun & Kindberg, 1892, p. 125. 

 

Pohlia nutans (Hedw.) Lindb. NY CHECK

 

CAN: As Webera nutans Hedw., Queen Victoria Niagara Falls Park .Cameron 1895.

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring); P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF); with Leptobryum pyriforme.

 

CAN: Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. W side; soil on clay, with Polytrichum commune. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Pohlia wahlenbergii (Web. & Mohr) Andr. NY CHECK

 

CAN: As Webera albicans Schimp., Queen Victoria Niagara Falls Park .Cameron 1895.

 

USA: Goat Island, south side, flats just west of Three Sisters, just west of Three Sisters, island margin of Cornus stolonifera (Eckel 1990b).

USA: Second Sister Island, east end, 2 Nov. 1988, Buck 16410 (BUF, NY).

Polytrichum commune Hedw. NY CHECK.

 

CAN: Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. W side; soil on clay, with Pohlia nutans. P. M. Eckel s.n. Sept. 21, 1998 (BUF); Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. Wet soil; on hummock. among Oaks. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Polytrichum formosum Hedw.

 

USA: New York, Erie Co.: Buckhorn Island, open swamp woods, Quercus rubra, Carya ovata, Acer rubrum, Ulmus americana: rather dry hummock in woods, Eckel July 26, 1998. The capsules were to 5 mm (not P. longisetum).

 

Polytrichum juniperinum Hedw. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.   

 

Polytrichum ohioense Ren. & Card.

 

CAN: Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Soil elevation, forest interior. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF).

 

Porotrichum alleghaniense (C.M.) Grout  = Thamnobryum alleghaniense (C. Muell.) Nieuwl. 

 

Pottia starkeana ssp. minutula (Schleich.ex Schwaegr.) Chamberl.

 

2005 Rare Plant Bryophyte Status List S1

 

USA: Eckel upper pavement at Devil's Hole Eckel 631685 (BUF), Eckel, l986. gorge of Niagara River, Devil's Hole, upper pavement in dolomite caprock overlooking the river, bare, sterile patches in lawn margins ‑ not in bare patches in the middle of the lawns.

 

USA: Lewiston. South of Lewiston, Devil's Hole State Park, rim of gorge of Niagara River, on bare soil in lawn near top of stairs into gorge. R. Zander & P. Eckel 8612282 Oct. 28, 1986 (BUF).

 

Pottia truncata (Hedw.) Fuernr. ex BSG var. truncata

 

CAN: Zander Foster's Flats, lower slopes

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Flat lawns above gorge, compact clay margins of exposed dolomite bedrock (var. major nearby)

With R. H. Zander.  P. M. Eckel  (Dupl. CANM 133585 April 5, 1985) (BUF).

 

 CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Barbula unguiculata, May 27, 1991, Eckel No. 9302250 G (BUF).

 

USA: Green Island on the Niagara R., just N of Goat Isl. Old dolomite ballast on south side, wet to moist river margin. With Barbula unguiculata, Phascum cuspidatum. PM Eckel, Marguerite P. Eckel 9004155a April 15 1990(BUF).

 

USA: Goat Island on the Niagara River. Soil at base of tree by the restaurant beside the driveway. With Barbula unguiculata P. M. Eckel  det. Zander 881031 Oct. 29, 1988 (BUF).

 

USA: Lewiston. Artpark, alvar on ledges at top of embankment of Niagara River beside stone piers of old Suspension Bridge, Syntrichia ruralis, Saxifraga virginiensis, Rumex acetosella. P. M. Eckel s.n. May 24, 2002 (BUF).

 

Pottia truncata var. major (Web. & Mohr) BSG

 

2005 Rare Plant Bryophyte Status List S1 [GeG5T 3T5 as Global Rank]

 

=Tortula modica Zander

 

CAN: Eckel‑Zander, Foster's Flats, above.

 

USA: halfway up talus slope, between steps & Whirlpool, Eckel 522386 BUF.

 

CAN: Ontario, City of Niagara Falls, on soil of slope above Table Rock House, next to inclined railway, rudimentary peristome present, spores mature, Zander & Eckel, April 4, 1993 (BUF). 

 

Pseudoleskeella tectorum (Funck ex Brid.) Kindb. ex Broth. NY CHECK

 

2005 Rare Plant Bryophyte Status List SU

 

As Leskea wollei Aust. "Niagara Falls. (Wolle.)," Macoun & Kindberg, 1892, p. 170. 

 

As Leskea wollei, Austin. 1874. Bull. Torr. Club, Vol.5:21‑24 (p22). "Niagara Falls (Wolle) ..." (Lesquereux & James, 1884, p. 304). A possible type locality. The description in the bulletin also included Lake Superior region and a collection by Prof. John Macoun.

 

Pterogonium trichomitrion, Drumm.

 

CAN: Drummond, Musci Amer. (Rocky Mts.) No. 78 (as Pterogonium trichomitrion Hedw.) "Hab. Trunks of trees in Upper Canada; about the Falls of Niagara." (NY). 

 

USA: Zander, Devil's Hole, lower slopes, Zander 3513 (BUF, NYS); Eckel l986.

 

CAN: As Leptodon trichomitrius, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

As Leptodon trichomitrius, "... about Niagara Falls. (Drummond.)," Macoun & Kindberg, 1892, p. 161. 

 

Ptychomitrium incurvum (Schwaegr.) Spruce

 

2005 Rare Plant Bryophyte Status List SH

 

CAN: As Grimmia hookeri nov. sp., Hab. On a stone near the Falls of Niagara in Upper Canada; rare. No. 61 (NY), Drummond; Musci Americani No. 61, 1828. 

 

"Ontario (Niagara Falls) and New York ..." Crum & Anderson p. 670. NY Checklist for 21,24,25 26 non 5.

 

CAN: As Pelychomitrium incurvum, Queen Victoria Niagara Falls Park .Cameron 1895. Probably one of Cameron's outrageous typographical errors.

 

Lesquereux & James (1884, p. 157) cite the substrate as "exposed rocks, especially sandstone," "Canada, near Niagara Falls (Drummond)."

 

As Grimmia Hookeri, Drumm."On a stone near the Falls of Niagara, in Ontario; rare. (Drummond.)," Macoun & Kindberg, 1892, p. 79. 

 

Pylaisia intricata (Hedw.) Grout

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.

 

Pylaisia polyantha (Hedw.) Grout

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.

 

CAN: As Pylaisia heteromalla Bruch et Schimp., Queen Victoria Niagara Falls Park Cameron 1895.

 

Rhodobryum ontariense (Kindb.) Paris in Kindb.

 

CAN: Eckel, Foster's Flats, halfway down.

 

CAN: As Bryum ontariense Kindb, Queen Victoria Niagara Falls Park . Cameron 1895.

 

Rhynchostegium serrulatum (Hedw.) Jaeg. & Sauerb. NY CHECK

 

USA: Gorge of Niagara River, "Upper Great Gorge" trail from Schoelkopf Museum towards Whirlpool. Lockport Dolomite. 1/3 way down. Soil. R. H. Zander 3759 Oct. 6, 1971 (BUF).

 

USA: Eckel, upper slopes near RR Bridge.

 

USA: Eckel, bluffs overlooking Lewiston, old growth Oak-Hickory   woods, with Weissia controversa (Eckel June 9, 1995) MO.

 

USA: Eckel between steps & Whirlpool.

 

Rhytidium rugosum (Hedw.) Kindb. NY CHECK

 

Hypnum rugosum Ehrh. ex Hedw.

 

USA; (As Hypnum rugosum Ehrh.) "Exposed places on rocks. Goat Island, Lesqx. Whirlpool wood, G. W. Clinton" (Peckl 1866).

 

(As Hypnum rugosum Ehrh.) (author given as Hedw.) Rare. Niagara Falls; Whirlpool woods, Day 1883

 

[No country] [as Hypnum rugosum] "Niagara Falls", Francis Wolle, 1873. ex herb. Coe Finch Austin (NY).

 

[No country] "Hab. Niagara Falls, 1873", Francis Wolle, ex herb. Eugene A. Rau (NY).

 

"... most characteristic of northern and montane flora" (Crum & Anderson, p. 1211).

 

Rhytidiadelphus triquetrus (Hedw.) Warnst.

 

CAN: As Hypnum trignetrum, Queen Victoria Niagara Falls Park .Cameron 1895. prob. H. triquetrum.

 

CAN: As Hylocomium triquetrum (Hedw). BSG "Roundhouse woods, Niagara Falls, 1901. (Macoun.)," Macoun, 1902, p. 318. 

 

Seligeria calcarea (Hedw.) BSG NY CHECK

 

2005 Rare Plant Bryophyte Status List SH

 

"The discovery of this rare and minute species in our [New York] State (as well as of [S. recurvata] is due to the earnest and faithful researches of G. W. Clinton (Peck 1866).

 

Seligeria calcarea Devil's Hole, Niagara Co. Day 1883.

 

USA: Devil's Hole, G. W. Clinton [two identical specimens mounted on cards] [NYS]

 

USA: Lewiston. Devil's Hole, Niagara. [Handwriting of Leo Lesquereux] [outside label in Clinton's handwriting G. W. Clinton legit s.n. s.d.] [1860-1870's] (BUF).

 

Seligeria campylopoda Kindb. ex Macoun & Kindb.

 

2005 Rare Plant Bryophyte Status List G3G5  S2S3

 

"Quite common and in fine fruit on damp limestone rocks, Foster's Flats, Niagara River, Ont., May 16th, 1901. (Macoun.)," Macoun, 1902, p. 206.

 

USA: Vitt, l976, New York: Buffalo, Clinton n.s. (MICH, MO, NY);

 

USA: Devil's Hole, Clinton, Oct., 1865 (FH).

 

CAN: "On limestone rocks, Niagara River," John Macoun, 11/5, 1901, New York Botanical Garden Mosses from Ontario, Canada. (No.4). c.fr. (NY)

 

USA: Devil's Hole, on stones, Oct. 24, 1866, Clinton [as Seligeria recurvata; mixed with Hymenostylium recurvirostrum] [NY]

 

USA: Devil's Hole, G. W. Clinton [as S. recurvata] [NYS]

 

CAN: Niagara on the Lake. Gorge of Niagara River, "First Gorge" on

escarpment at Redan Battery Site on moist rock. det. R. H. Zander 1995

R. H. Zander 3708 Sept. 29, 1971 (BUF).

 

USA: Upper path lower Arch Bridge, Eckel 2121885 (ALTA, BUF), (Eckel l986a).

 

USA: Eckel between steps & Whirlpool 521386 BUF

 

Seligeria pusilla (Hedw.) BSG

 

"Devil's Hole, near Niagara Falls (G. W.  Clinton) ..." (Lesquereux & James, 1884, p.96‑97). This species was also reported on "Shaded limestone rocks, St. Louis, Missouri (Drummond) … New Jersey (Austin); Kelly's Island, Lake Erie (Lesquereux)."

 

Seligeria recurvata (Hedw.) BSG NY CHECK

 

2005 Rare Plant Bryophyte Status List S2S3

 

USA: Rocks in ravines. Devil's hole, Niagara county … , G. W. Clinton (Peck 1866).

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.  

 

USA:  Seligeria recurvata Devil's Hole, Niagara River,  Charles Peck in Day 1883.

 

USA: (as S. recurvata var. arcuata) "Devil's Hole, Niagara (Clinton, Mrs. Roy) …." (Lesquereux & James 1884).

 

USA: New York: a specimen of George Clinton's, with no locality was sent to Elizabeth Gertrude Britton by Smith Ely Jecliffe [?, sp.] on January 23, 1892 on which she had written "probably Niagara Falls" by her initials (NY).

 

USA: Vitt, l976, New York: Niagara Co., Devil's Hole, Clinton 24 (NY); Goranda (?) [Gowanda] Cuttasanya [Cattaraugus] Creek, "coll. Fr. H.V.& Fr. Ph. B." (MO).

 

USA: As var. arcuata, previously called Weisia Seligeri, Hook & Wils. in Drumm., "Devil's Hole, Niagara (Clinton, Mrs. Roy)..." (Lesquereux & James, 1884, p. 97). 

 

USA: "That little moss sent with letter of Sept. 21 proves to be a very fine and rare species, viz: Seligeria recurvata Br. Eur. Mr. Sullivant has described it in his manual or rather in Gray's Manual 2nd 3d. but I do not know from what specimens for it is the first time that I see it in America. It is a very good addition to your catalogue of New York mosses. I wish you had found it in plenty. I would have put it in the Musci. I have no doubt that you may find at the same place S. tristicha which generally grows with it. It is still shorter and blackens by its foliage the reversed surface of wet overhanging rocks. Whenever you go to Devil's Hole, please look for it carefully. Mr. Sullivant has found it around Columbus in a kind of grotto limestone which has been destroyed long time ago by the process of quarrying limestone. I could never find it again." Leo Lesquereux to George Clinton, Oct.  17th, 1865 (vol.2:10)

 

Sphagnum imbricatum Hornsch. ex Russ.

 

"... in damp woods, near the roundhouse, Niagara Falls, Ont., May 14th, 1901. (Macoun.)," Macoun, 1902, p. 183.

 

Sphagnum palustre L.

 

Sphagnum cymbifolium (Ehrh.) Hedw.

 

CAN: As Sphagnum cymbifolium, Queen Victoria Niagara Falls Park .Cameron 1895.

 

Sphagnum nemoreum Scop.

 

Sphagnum acutifolium Schrad.

 

CAN: As Sphagnum acutifolium, Queen Victoria Niagara Falls Park .Cameron 1895

 

Note that these two localities are probably also the stations for the bog plants that have been reported for the Niagara Parks area.

 

Taxiphyllum deplanatum (Bruch & Schimp. ex Sull.) Fleisch.

USA: First Sister Island, western half, 1 Nov. 1988, Buck 16345 (BUF, NY), 2 Nov. 1988, Buck 16426 (BUF, NY).

Tetraphis pellucida Hedw. NY CHECK

 

USA: Second of the Three Sisters Islands in the Niagara River just S of Goat Island. Muck alluvium, west end, with Didymodon fallax, humectated tree roots. P.M.. Eckel 881103 Oct. 29, 1988 (BUF); Second Sister Island, west end, tree roots, wet in high water, Oct. 29, 1988 (BUF); west end, on rotten log, 2 Nov. 1988, Buck 16371 (BUF, NY).

 

USA: Goat Island State Park, oak-maple forest over limestone; Three Sisters Islands, west end of Second Sister. William R. Buck 16371 Nov. 2, 1988 (BUF, NY).

 

CAN: Navy Island in the Niagara River, just south of Niagara Falls, northwest and northern river margins. Stumps, west side. Didymodon fallax. P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF).

 

Thamnium alleghaniense, see Thamnobryum alleghaniense (C. Muell.) Nieuwl.

 

Thamnobryum alleghaniense (C. Muell.) Nieuwl. NY CHECK

 

=Hypnum alleghaniense C.Muell.

 

=Porotrichum alleghaniense (C.M.) Grout

 

USA: (As Hypnum alleghaniense, C. Mull. [sic]) "Moist rocks along streams and clefts of rocks on mountains. … Devil's Hole, G. W. Clinton." (Peck 1866),

 

USA: (As Hypnum alleghaniense). Rare. Devil's Hole, Niagara River. Day 1883.

 

CAN: (As Hypnum alleghaniense), Queen Victoria Niagara Falls Park .Cameron, l895. 

 

CAN: (As Thamnium alleghaniense), Queen Victoria Niagara Falls Park .Cameron 1895.

 

USA: New York: Devil's Hole, G. W. Clinton (NYS) [glued on card, "1"]

 

USA: Clinton specimen for Devil's Hole under Hypnum alleghaniense C. Muell. herb no. 38733 (BUF).

 

USA: Lewiston. Devil's Hole. Niagara River-Valley near Buffalo.  Limestone on rocks in dry creek valley. R. Duell 805 May 19, 1981 (BUF).

 

Thelia hirtella (Hedw.) Sull. in Sull. & Lesq.

USA: First Sister Island, western half, Eckel 880727, July 8, 1987 (BUF).

Thuidium abietinum (Hedw.) BSG.

= Abietinella abietina (Hedw.) Fleisch.

= Hypnum abietinum Hedw.

 

USA: "Rocks. Goat Island, Lesqx. Whirlpool wood, Niagara Falls, G. W. Clinton." (Peck 1866).
 

[No country] Niagara Falls, Francis Wolle, 1873; ex herb. Coe Finch Austin (NY). 

 

[No country] Hab. Niagara Falls, Aug. 1873, Francis Wolle, ex herb. Eugene A. Rau, Bethlehem, Pa. (NY).

 

USA: New York. Niagara Falls, N.Y. July 1850. "7". Thomas P. James, collector, No. 7. Mosses of Ezra Michener Herbarium, received from C. L. Shear, June 1930 (US).

 

USA: Musci Boreali‑Americani [first edition], W. S. Sullivant et L. Lesquereux, MDCCCLVI, No. 280. Hab. ... "in insula Goat Island dicta juxta cataractam Niagarae" (NY). 

 

USA: No. 413 [second edition Musci Boreali‑Americani] ex herb. Leo Lesquereux, "... in insula Goat Island dicta juxta cataractam Niagarae" (NY, also NYS). 

 

CAN: "On dry limestone rocks, Queenston Heights, Ont., May, 1901. (Macoun.)," Macoun, 1902, p. 284.

 

CAN: Queen Victoria Niagara Falls Park .Cameron 1895.

 

CAN: Niagara Glen (Wintergreen Flats), top dolomite surface of cliff overlooking gorge, with Tortella tortuosa, Eckel 9302091a, May 6, 1986 (BUF).

 

USA: Lewiston. Artpark, alvar on ledges at top of embankment of Niagara River beside stone piers of old Suspension Bridge, Syntrichia ruralis, Saxifraga virginiensis, Rumex acetosella. P. M. Eckel s.n. May 24, 2002(BUF).

 

"A calciphile, on dry, exposed rocks, soil, or turf, on sand of partially stabilized dunes, among talus at the base of cliffs, or on humus in open, coniferous stands" Crum & Anderson p. 911.

 

Thuidium delicatulum (Hedw.) BSG

 

CAN: Queen Victoria Niagara Falls Park . Cameron 1895.

 

USA: as Hypnum delicatulum L. Devil's Hole, Clinton specimen 38757.

 

USA: (var. delicatulum) Second of the Three Sisters Islands in the Niagara River just S of Goat Island. Muck alluvium, west end, with Didymodon fallax, humectated tree roots. P.M. Eckel 881105 Oct. 29, 1988 (BUF).

 

USA: Goat Island State Park, oak-maple forest over limestone; Three Sisters Islands, west end of Second Sister. William R. Buck 16373 Nov. 2, 1988 (BUF, NY).

 

CAN: (var. delicatulum) Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Dominant on soil, island west side; Beech-Maple, Carya-Quercus forest; Atrichum undulatum. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Shaded soil; shore; top of embankment; + Amblystegium varium; Barbula unguiculata; Anomodon rugelii. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

Thuidium pygmaeum BSG NY CHECK

 

2005 Rare Plant Bryophyte Status List  S1

 

Hypnum pygmaeum (BSG) Sull. et Lesq. is this species.

 

Note that Thuidium minutulum (Hedw.) BSG, a moss more closely associated with decayed wood and the bases of trees  may be confused with T. pygmaeum, which is a calcifile. Its habitat is on rock and has papillose stems and branches. T. minutulum is more northerly in distribution.

 

CAN: "Rocks. Foster's Flat, G. W. Clinton. Rare." (Peck 1866). Since the Peck catalogue was for New York State it appears Peck was under the impression Clinton's "Foster's Flat" was in USA.

 

CAN: (as Hypnum pygmaeum (BSG) Sull. et Lesq.) Rare. Foster's Flat, Niagara River, Ont., Day 1883. 

USA: Goat Island, central woods; thin soil on limestone cobbles (Eckel 1990b).

 CAN: Clinton specimen Foster's Flat [May?] 4, herb no 38777 (BUF).

 

USA: Niagara (American) Falls, N.Y. on moist limestone and on small limestone pebbles. A. T. Beals, 26 Aug., 1917 (NY).

 

USA: Niagara. On moist limestone and on small limestone pebbles. A. T. Beals, Aug. 26, 1917 (NY). 

 

CAN: Foster's Flat. G. W. Clinton [glued on card] (NYS).

 

CAN: Niagara Glen, just N of Whirlpool. Base of boulder, dolomite, N-facing aspect. With Lophocolea heterophylla. P.M.Eckel 231986 April 11, 1985 (BUF).

 

Thuidium recognitum (Hedw.) Lindb.

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: New York, Erie Co., Grand Island Twp.,  Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring); P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF); with Mnium cuspidatum.

 

USA: First of the Three Sisters Islands off Goat Island in the Niagara River. Dolomite boulder top. West end. P.M.Eckel 8712251 April 26, 1987 (BUF); Goat Island State Park, oak-maple forest over limestone; Three Sisters Islands, west end of First Sister. William R. Buck 16343a Nov. 1, 1988  (BUF, NY).

 

 

Timmia megapolitana Hedw. NY CHECK

 

USA: Zander, Ongiara, lower wet Zander 3521b, (BUF), Eckel l986.

 

USA: Devil's Hole upper Eckel 931685 (BUF, NYS), Eckel l986.

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Moist, vertical dolomite block in late snow melt, with Conocephalum conicum. With R. H. Zander P. M. Eckel 23885 April 5, 1985 (BUF).

USA: First Sister Island, western half, Eckel 880719, July 8, 1987 (BUF).

Timmia megapolitana var. bavarica (Hessl.) Brid.

 

As Timmia bavarica Hessler.  "On limestone rocks, Foster's Flats, Niagara Falls, 1901. (Macoun.)," Macoun, 1902, p. 262. 

 

Tortella fragilis (Drumm.) Limpr. NY CHECK

 

[?] Niagara Falls, Coe Finch Austin, July, 1874 (NYS) [no country indicated on note inside packet: later prepared external packet indicates New York].

 

CAN: Table Rock, Niagara Falls, Herb. Coe Finch Austin, July, 1874 (NY).

 

USA: Goat Island, Niagara Falls, N.Y. Ground, June 1, A. B. [A. Barroni?] ca. 1885 [herb. purchased by Columbia College 1885‑1887]

 

USA: Foot of Goat Island, just outside spray area of Horseshoe Falls, boulder, thin soil, mid-slope. With Hyophila involuta, Hymenostylium recurvirostrum, Trichostomum crispulum. R. H. Zander 3484b Oct 28, 1970 (BUF).

 

Tortella humilis (Hedw.) Jenn.

 

=Barbula caespitosa Schwaegr.

 

USA: gorge of Niagara River, near bottom of slope just north of sewage plant, on thin soil over calcareous rock. R. Zander & P. M. Eckel 8511211 Nov. 17, 1985 (BUF).

USA: First Sister Island, western half, 1 Nov. 1988, Buck 16347 (NY).

Tortella inclinata (Hedw. f.) Limpr.

 

=Barbula inclinatula, C. M. & Kindb. in Mac. & Kindb.

 

As Barbula inclinatula "Sterile specimens collected on limestone rocks along the Niagara River, near Queenston, Ont., May 13th, 1901, are doubtfully refered here. (Macoun.)," Macoun, 1902, p. 212. 

 

Ireland and Cain (1975) report this species only from the Bruce Peninsula in Ontario.

 

Tortella tortuosa (Hedw.) Limpr.

 

=Barbula tortuosa, (Hedw.) Web. & Mohr.

 

=Tortula tortuosa, Ehrh. (Lesquereux & James, 1884).

 

USA: (as Barbula tortuosa) "Rocks. Goat Island, Lesqux. Common about Niagara Falls, G. W. Clinton." (Peck 1866).

 

USA: "Goat Island, Niagara Falls..." (Lesquereux & James, 1884, p. 129). 

 

CAN: as Barbula tortuosa Web. and Mohr) Niagara Falls, Day 1883. 

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.

 

Podpera (1954) cited a specimen from "Niagara."

 

USA: Musci Boreali‑Americani, W. S. Sullivant et L. Lesquereux, MDCCCLVI, No. 97b [first edition], "Hab. ad rupes calcarias insulae Goat Island dictae, juxta cataractam Niagarae" (NY.) [Sullivant & Lesquereux, Musci Bor.‑Amer., ed 1, 97b, and ed. 2, 137 (as Barbula):specimen at CANM with same Latin, numbered no. 137

 

USA: Niagara Falls, G. W. Clinton (NYS) [internal label by Peck: 1 (Niagara) & 2 (Luzerne), G. W. Clinton, 3 (Mt. Marcy) & 4 (Helderberg Mts.) C. H. Peck, "Coll. N.Y. State"]. 

 

USA: Musci Boreali‑Americani, W. S. Sullivant et L. Lesquereux, Editio Secunda, 1865, ad rupes calcarias insulae Goat Island dictae, juxta cataractam Niagarae, same collection as above (NY).

 

USA: (as Barbula tortuosa Web. et Mohr) Hab. ad rupes calcarias insulae Goat Island dictae, juxta cataractam Niagarae. (NYS) [Sullivant & Lesquereus, Musci Bor.‑Amer., ed 2, No. 137] (BUF).

 

USA: (as Barbula tortuosa Web. et Mohr.)City of Niagara Falls On the ground.[hand writte label] Thomas P. James s.n. July 1860 (BUF).

 

USA: Niagara Falls, N.Y., G. W. Clinton, ex Coll. Charles H. Peck, Albany, NY (NY)

 

USA: Goat Island, Niagara Falls, G. W. Clinton, June, 1865, Herb. Coe Finch Austin (NY).

 

USA: First Sister Island, Niagara Falls, AAAS, E.G. B & E. A. Rau, Aug. 22d, 1886 (NY, meeting of the American Association for the Advancement of Science).

USA: Tortella tortuosa (identified as Barbula fragilis). Niagara Falls, N.Y. "Sister Id., no. 1." August 22d, 1886. AAAS Mg. E[lizabeth] G B [Britton] & E[ugene]. A. Rau New York Botanical Garden [collected during the 1886 meeting of the American Association for the Advancement of Science.]

[No country]: Niagara Falls, July 18, 1874, herb. Coe Finch Austin (NY) 

 

[inside packet "Niagara River, below the falls, July 18, 1874, duplicate J.B.S [L.?]"].

 

[No country]: Niagara Falls, Rev. Francis Wolle, 1873, Herb. Coe Finch Austin (NY).

 

USA: Zander, Whirlpool, upper slopes.

 

CAN: Niagara Glen, just N of Whirlpool. Moist thin soil in depressions in ls boulders, Acer - Tsuga canadensis wood. W. Brotherella recurvans. P.M.Eckel NF17 Feb. 19, 1983 (BUF); Niagara Glen, just N of Whirlpool. Thin soil over limestone boulder, shade. With Anomodon rostratus, Encalypta procera, Platydictya confervoides. With R.H.Zander. P.M.Eckel NF1 Feb. 19, 1983 (BUF); Niagara Glen (Foster's Flats) on the Niagara River. Frequent on moist boulders. With R. H. Zander. P. M. Eckel 13585 April 5, 1985 (BUF); Niagara River gorge, Niagara Glen, limestone. R. H. Zander & P. M. Eckel 4724 Mar. 4, 1979 (BUF).

 

USA: Between Devil's Hole and Whirlpool steps, Eckel 614486.

 

USA: Boulder field, halfway down Whirlpool steps, with Anomodon viticulosus, Bryum lisae var. cuspidatum, Campylium chrysophyllum, Gymnostomum aeruginosum, Marchantia polymorpha, Encalypta procera, May 6, 1986, Eckel 9302092f (BUF). 

 

USA: north of Whirlpool State Park, just N of DeVeaux stone steps down calcareous gorge face, moist shaded wooded bottom above riverbank on moist boulders, with Brachythecium oxycladon, B. curtum, Eckel, Sept. 23, 1991 (BUF).

 

 CAN: Niagara Glen (Wintergreen Flats), top dolomite surface of cliff overlooking gorge, with Thuidium abietinella, Eckel 9302091b, May 6, 1986 (BUF) . 

 

USA: First of the Three Sisters Islands just S of Goat Island in the Niagara River. West end, dolomite boulder, w. Bryoeryth. recurv., Timmia megapolitana, Anomodon attenuatus, etc. P. M. Eckel 880728 July 8, 1987 (BUF).

 

Tortula mucronifolia Schwaegr. NY CHECK

 

USA: (As Barbula mucronifolia Schwaegr.) "Stones; Devil's Hole and Portage, G. W. Clinton." (Peck 1866).

 

USA: (As Barbula mucronifolia Schwaegr.) Devil's Hole, Niagara River, Day 1883.

 

USA: North American Mosses, Eugene A. Rau, Bethlehem, Pa., Hab. banks of Luna Island, Niagara Falls, Aug., 1886 (NY), c. fr.

 

USA: Niagara Falls, Rev. Francis Wolle, 1873, c. fr. (NY).

USA: (as Barbula mucronifolia) Luna Island, Niagara Falls, N.Y. E. G. Britton & E. A. Rau. Aug. 21, 1886, North American Mosses named and presented by E. G. Britton, Aug. 21, 1886.

USA: (as Barbula mucronifolia) Hab. banks of Luna Island, Niagara Falls, North American Mosses, Aug. 1886, Eugene A. Rau, Bethlehem, Pa. [printed label] (NY).

USA: (as Barbula mucronifolia) Hab. banks of Luna Island, Niagara Falls, North American Mosses [printed label], August 1886 Eugene A. Rau, Bethlehem, Pa. [printed label] [New York Botanical Garden] (NY).

USA: Luna Island, Niagara Falls, E. G. Brisson, E. A. Rau: AAAS Meeting, Aug. 21, 1886 (NY), c.fr. [= American Association for the Advancement of  Science].

 

USA: Zander, Ongiara, lower slopes; Ongiara Gorge, between the (S) Whirlpool and the stone steps (N) descending to the Niagara River. With Didymodon rigidulus, Encalypta. P.M.Eckel 622386 Mar. 21, l986 (BUF).

 

USA: Eckel between steps Whirlpool 821386 BUF.

 

USA: Luna Island, 1 Nov. 1988, Buck 16329 (BUF, NY).

USA: First Sister Island, western half, Eckel 880718, July 8, 1987 (BUF).

USA: First Sister Island, eastern half,  Eckel 880708, July 8, 1988 (BUF).

USA: Third Sister Island, west end, in small solution vugs in dolomite, P. M. Eckel 8612338, April 27, 1986 (BUF).

USA: Green Island on the Niagara River, just No of Goat Island. Old dolomite ballast on south side, wet to moist river margin. With Pottia truncata, Phascum cuspidatum. P.M. Eckel, Marguerite P. Eckel 9004156 April 15 1990 (BUF).

 

CAN: Niagara Glen, N of the Whirlpool of the Niagara River. In lawns above the crest, dolomite exposures: on edge of the soil mat. P. M. Eckel 8612337 May 6, l986 (BUF).

 

CAN: Niagara Glen (Foster's Flats). On rock, halfway down to Niagara River, dolomite boulder field. With Didymodon rigidulus, Fissidens bryoides. P.M.Eckel 1074901 April 11, 1985 (BUF).

 

Tortula ruralis (Hedw.) Gaertn., Meyer & Scherb.

 

CAN: As Barbula ruralis Hedw, Queen Victoria Niagara Falls Park.Cameron 1895; Macoun (p. 58, 1892) cited a specimen of Cameron's from Niagara Falls.

 

USA: First of Three Sisters Eckel 19786 BUF, NYS (Eckel, l986b).

 

CAN: Dufferin Islands, mats on soil, Eckel & Zander 9304221 (BUF).

 

USA: Lewiston. Artpark, alvar on ledges at top of embankment of Niagara River beside stone piers of old Suspension Bridge, Ceratodon purpureus, Saxifraga virginiensis, Rumex acetosella. P. M. Eckel s.n. May 24, 2002(BUF).

 

USA: Gorge of Niagara River, below Schoelkopf Museum, on limestone about 1/5 way up from river. Richard H. Zander & P. Eckel 8704231 April 16, 1987(BUF).

 

Tortula subulata Hedw.

 

Barbula subulata (Hedw.) P. Beauv.

 

Given as questionable in Ontario, lit for YT and positive for BC in Ireland et al mosses of Canada, 1980

 

As Barbula subulata (Hedw.) P.  Beauv. "...  on earth on rock, Niagara River, Ont. (Macoun.)," Macoun, 1902, p.  214.

 

This western species intergrades with Tortula mucronifolia in the west (Lawton, 1971).  Macoun's specimens from Niagara are probably T. mucronifolia. 

 

Trichostomum crispulum Bruch in F. A. Muell.

 

Trichostomum crispulum has remained ignored for the area N of Mexico, probably due to its confusion with related taxa and Weisia species, there being no fruiting material.  However, according to Zander (study in preparation), this species grows abundantly throughout Mexico and is indeed Trichostomum crispulum, with typical cuculate apex, etc. This species may be new to eastern North America

 

USA: Goat Island. Central Acer‑Fraxinus mixed old woods, soil, west end of woods in spray zone of the Horseshoe Falls. Sandy loam. Eckel & Eckel, April 15, 1990, No. 9004273 (BUF). 

 

USA: Foot of Goat Island, just outside spray area of Horseshoe Falls, boulder, thin soil, mid‑slope, with Hyophila involuta, Hymenostylium recurvirostrum, Tortella fragils, R. H. Zander 3484a, Oct. 28, 1970 (BUF). 

USA: Trichostomum crispulum. Just outside spray area of Horseshoe Falls, boulder, thin soil, midslope, with Hyophila involuta, Hymenostylium recurvirostrum, Tortella fragilis, Zander 3484a Oct. 28, 1979 (BUF).

Weisia brachycarpa (Nees & Hornsch.) Jur.

 

As Weissia hedwigii  2005 Rare Plant Bryophyte Status List S1

 

CAN: City of Niagara Falls, base of wooded embankment overlooking the horseshoe Falls. In spray on calcareous clay in lawn, May 27, 1991 P. M. Eckel no. 9302250 (BUF), with Barbula unguiculata, Amblystegium varium Physcomitrium turbinatum, Fissidens taxifolius. 

 

Weisia controversa Hedw. NY CHECK

 

USA: Upper great gorge. S of sewerplant, Eckel 29786 (BUF)

 

USA:  bluffs overlooking Lewiston, dry Oak-Hickory old growth, on soil with Rhynchostegium serrulatum (Eckel June 9, 1995) MO, c. fr. deoperculating.

USA: Luna Island, 1 Nov. 1988, William R. Buck 16315 (BUF, NY).

USA: First Sister Island, western half, small population, dolomite boulder, with Tortula mucronifolia, synoicous, 8706070, 1986; 1 Nov. 1988, William R. Buck 16340 (BUF, NY).