Preliminary Cryptogamic (Moss, Lichen and Liverwort) Flora of the Canadian and American Gorge at Niagara Falls 1. Mosses

P. M. Eckel
Res Botanica, a Missouri Botanical Garden Web site

http://www.mobot.org/plantscience/ResBot/index.htm

June 21, 2004

 

PRELIMINARY CRYPTOGAMIC (MOSS, LICHEN AND LIVERWORT) FLORA OF THE CANADIAN AND AMERICAN GORGE AT NIAGARA FALLS

1. Mosses

 

P. M. Eckel, Missouri Botanical Garden, PO Box 299, St. Louis, MO 63166-0299 USA

 

 

 

Grimmia alpicola Hedw.

 

USA: Eckel below between steps and Whirlpool 422386 BUF

 

USA: [var. alpicola] Devil's Hole, near bottom with Bryum capillare var. flaccidum, Entodon seductrix, Bryhnia graminicolor, May 21, 1988, Eckel No. 9303181C (BUF). 

 

USA: Devil's Hole, upper slopes, Eckel 226386.

 

USA: (var. alpicola) Goat Island on the Niagara River, wooded slope just N of first 3-Sisters Islds Bridge, on dolomite boulder. Plants to over 7mm, lvs to 2mm, capsules immersed. P. M. Eckel 881003 Sept. 29, 1988 (BUF).

 

USA: (var. alpicola) Goat Island on the Niagara River, in depression on dolomite block of first bridge to the Three Sisters Islands. Plants to over 7mm, lvs to 2mm, capsules immersed. P. M. Eckel 881002 Sept. 29, 1988 (BUF).

USA: (As Schistidium alpicola). Luna Island, 1 Nov. 1988, Buck 16319 (NY).

USA: Green Island on the Niagara R just N of Goat Island. Old dolomite ballast on S side, occasionally submersed, wet to moist river margin. PM Eckel, Marguerite Eckel 9004158 April 15, 1990 (BUF).

USA: (As Schistidium alpicola): Second Sister Island, east end, 2 Nov. 1988, Buck 16411 (NY).

USA: var. alpicola. Dolomite bridge, First Sister Island,  in depression, with lichens, Eckel, Sept. 29, 1988 (BUF).

Grimmia alpicola var. rivularis (Brid.) Wahlenb.

 

CAN: Niagara Glen, just N of Whirlpool. Moist thin soil in depressions in limestone boulders. w. Tortella tortuosa, Didymodon rigidulus. With R.H.Zander. P.M.Eckel NF17 Feb. 19, 1983 (BUF).

 

Grimmia apocarpa Hedw. NY CHECK as var. apocarpa

 

USA: Zander, Devil's Hole lower area between Devil's and whirlp on USA side.

 

CAN: Grimmia apocarpa, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

As Grimmia apocarpa var. stricta, Drumm. Musc. Bor. Am. No. 55 by Macoun & Kindberg:] "... Niagara Falls (Macoun.)," Macoun & Kindberg, 1892, p. 62‑63. 

 

Grimmia calyptrata A doubtful record.

 

A western species collected by Drummond in the Niagara gorge in BUF is misplaced from collections he probably made out west. It is annotated as a new sp. Herb. No. 56168, . "On stones, Niagara, rare" annotated as Grimmia calyptrata Hook. on packet.  cf. Wils., Hook. J. Bot. This specimen is actually Grimmia calyptrate but misplaced, sec. Zander.

 

Grimmia pilifera P. Beauv.

 

2005 Rare Plant Bryophyte Status List  S3? [sic]

 

Grimmia pennsylvanica Schwaegr. ex Sull. & Lesq.

 

CAN: "near the Falls of Niagara; upon rocks (as Grimmia pennsylvanica), Drummond, Musci Americani No. 56 (NY). 

 

CAN: As Grimmia pennsylvanica, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN:  Niagara Glen (Foster's Flats) on the Niagara River. Dry, occasional seepage in dolomite caprock, upper path, with Didymodon rigidulus, Gymnostomum aeruginosum. With R. Zander. P. M. Eckel 2231185 April 5, 1985 (BUF).

 

Gymnostomum aeruginosum Sm. NY CHECK

 

Gymnostomum calcareum, Nees, Hornsch. & Sturm

 

Gymnostomum rupestre Schleich. ex Schwaegr.

 

[USA?] On rocks, Whirlpool, Niagara Falls. G. W. Clinton (with Seligeria sp., label indicates S. recurvata, but removed from packet) (NY).

 

CAN: As Gymnostomum rupestre Schleich. following Crum & Andeson p. 283, Queen Victoria Niagara Falls Park.  Cameron 1895

 

CAN: As Gymnostomum calcareum, Nees & Hornsch. [G. tortile, Drummond. Musc. Bor. Am. 23]

 

CAN: "... on calcareous rocks near the Whirlpool, Niagara Falls. (Macoun.)," Macoun & Kindberg, 1892, p. 12.

 

CAN: As Gymnostomum rupestre, Schwaegr., "... near the Whirlpool, Niagara Falls (Macoun.),"   Macoun & Kindberg, 1892 p. 13. 

 

CAN: Macoun & Kindberg (p. 13, 1892) reported (as G. rupestre) this species growing "near the Whirlpool, Niagara Falls ... (Macoun)."

 

CAN: As Gymnostomum calcareum NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: (as Gymnostomum rupestre) Hab. on ground towards 3 sister islands, Niagara Falls, Aug. 21, 1886, c. fr., Eugene A. Rau, Bethlehem, Pa. (NY)

 

USA: On Three Sisters Island, Niagara Falls, N.Y., Aug. 21, 1886, c. fr., AAAS meeting, Elizabeth Gertrude Britton (NY).

 

A specimen from NY determined as Barbula convoluta was identified by R. Zander (1996) to be Bryoerythrophyllum recurvirostrum mixed with Gymnostomum aeruginosum. The plants were collected at "Niagara Falls" by Wolle in 1873. There is a packet with Wolle's autograph on it, and the packet was constructed out of the fragment of a letter, presumably to Wolle, by Eugene A. Rau; the letter is unreadable due to its fragmentary nature.  Crum & Anderson (l981) discuss the occurrence of this species in North America and believe specimens identified as G. calcareum in eastern North America are in fact G. aeruginosum based on a variety of characters, p.  285. Zander, Bryologist 80:233‑69, 1977, synonymized G. calcareum with G.  aeruginosum.  Podpera (1954) reported a specimen of Gymnostomum calcareum from "Niagara." Macoun & Kindberg (p. 12, 1892) reported Gymnostomum calcareum "on calcareous rocks near the Whirlpool Niagara Falls.  (Macoun). 

 

USA: boulder field, halfway down Whirlpool steps, with Anomodon viticulosus, Bryum lisae var. cuspidatum, Tortella tortuosa, Campylium chrysophyllum, Encalypta procera, Plagiopus oederiana, Mnium thomsonii, Marchantia polymorpha, May 6, 1987, Eckel 9302092b (BUF); DeVeaux (Whirlpool) stone steps down calcareous gorge face, moist shaded wooded bottom above Niagara riverbank, on moist boulders, c. fr., with Bryoerythrophyllum recurvirostrum. Eckel Oct. 4, 1991 (BUF).

 

USA: Zander, Devil's Hole waterfall.

 

USA: Zander, Ongiara, lower slopes

 

USA: Zander, Prospect Park, lower slopes.

 

USA: At Niagara Falls, base of Goat Island, just below lower entrance to elevator, about 70 feet above waterline, in vug in large calcareous boulder. R. Zander, P. Eckel, W. Buck, R. Harris 8811012 Nov. 1, 1988 (BUF).

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Upper path, dolomite caprock, with Didymodon rigidulus, Grimmia pilifera. With R. Zander. P. M. Eckel 2131185 April 5, 1985 (BUF).

 

CAN: Above Queenston, on the escarpment just S of Brock's Monument on the gorge lip: Locust" Grove picnic area. Thin soil over moist rock" in stream bed. P. M. Eckel 232085 April 20, 1985 (BUF).

 

Haplocladium microphyllum (Hedw.) Broth. NY CHECK

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring). P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

CAN: (Welland Co., Bertie Twp.) Niagara River, Navy Island, 43*03'N, 79*01'W, just S of Niagara Falls; west side, bark base of tree; Oak woodland. Dicranum montanum, Ceratodon purpureus.  P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Hedwigia ciliata (Hedw.) P.‑Beauv. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron, 1895.

 

CAN: As Hedwigia albicans Lindb.  "... on rocks, Whirlpool Rapids, Niagara River, 1901. (Macoun.)," Macoun, 1902, p. 225. 

 

USA: Second Sister Island, west end, on glacial erratic, 2 Nov. 1988, Buck 16380 (NY).

 

CAN: (Welland Co., Bertie Twp.), Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. With Hypnum pallescens on granite glacial erratic; Fraxinus-Carya woods P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Helodium paludosum (Sull.) Aust. 

 

As Elodium paludosum Aust. "... Queenston and "roundhouse woods," near Niagara Falls, Ont., 1901. (Macoun.)," Macoun, 1902, p. 286. 

 

USA: Erie Co. Grand Island. Ubiquitous on Grand Island. Niagara R. Ann Glowny 1338 3-28-67 (BUF).

 

CAN: Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Forest interior; moist Red Maple forest. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Muck soil, with Climacium americanum P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

Herzogiella striatella (Brid.) Iwats.

 

Plagiothecium muehlenbeckii BSG.

 

CAN: "... on old logs at Foster's Flats, Niagara River, Ont., 1901. (Macoun.)," Macoun, 1902, p. 299. 

 

Homalotheciella cubcapillata (Hedw.) Broth.

 

2005 Rare Plant Bryophyte Status List. SH

 

As Homalothecium subcapillatum Schp. Devil's Hole, Niagara River. Day 1883.

 

Homomallium adnatum (Hedw.) Broth.  NY CHECK

 

USA: Devil's Hole, upper slopes, Eckel 726386  (BUF); (Eckel, l986b).

 

CAN: Niagara Glen (Foster's Flats). Boulder field. Base of dolomite block, shaded woodland, with Didymodon rigidulus Bryum capillare. P.M.Eckel 229486 April 11, 1985 (BUF)

USA: First Sister Island, western half, 1 Nov. 1988, Buck 16342 (NY).

USA: Second Sister Island, west end, on glacial erratic, 2 Nov. 1988, Buck 16364 (BUF, NY).

Hylocomium brevirostre (Brid.) Fleisch. NY CHECK

= Hypnum brevirostre (Brid.) Ehrh. 

 

Hylocomium splendens (Hedw.) BSG

 

Hypnum splendens Hedw.

 

CAN: As Hypnum splendens, Queen Victoria Niagara Falls Park.  Cameron 1895.

 

Hymenostylium recurvirostrum (Hedw.) Dix.

 

= Gymnostomum recurvirostrum Hedw.

On August 8, 1865, George Clinton recorded in his journal that this moss was "everywhere common on wet rocks" below the Biddle Staircase (on Goat Island, west facing face of the Niagara River gorge)..

[USA?] Hab. Niagara Falls, July 18, 1874, Coe Finch Austin, leg. with Didymodon tophaceus (NY).

 

USA: Three Sisters Islands, Niagara River, Aug. 1886. L.M.U.  [Underwood] (NY) with bistratose leaf margins in patches. 

 

USA: Hab. ad cataractam Niagarae. Musci Alleghanienses, W. S. Sullivant, 1845, No. 193 (As Gymnostomum curvirostrum Hedw.), (NY).

 

USA: Rocks. Niagara Falls, N.Y., July, 1865, c.fr., G. W. Clinton, det. Charles H. Peck (NY).

 

USA: Biddle's Stairs, on damp rocks at Niagara Falls below Biddle's Stairs, Goat Island, Sept. 1853, c. fr., Chapman collection (NY)

 

USA: Hab. in rupibus irriguis Niagarae cataractae. Sullivant & Lesquereux, Musci Bor.‑Amer. ed 1, No. 41 (As Gymnostomum curvirostrum Hedw.)(NY). This collection has capsules which differ from the typical variety by the "capsula ovato‑cylindrica fusco‑lutea," but is otherwise indistinguishable.

 

USA: Hab. in rupibus irriguis Niagarae cataractae. Sullivant & Lesquereux, Musci Bor. ‑Amer. ed. 2, 1865, No. 53 (NY). The NY specimen from Jaeger's Herbarium (No. 777) was annotated as var.  ovatum Sull. & Lesq. It is still "capsula ovato‑cylindrica fusco‑lutea". The specimen examined above as No. 41 of the first edition represented a population of medium to small stature. In No 53, two specimens, one tall, relative to the other, are glued to the card, the lesser probably the var. inferior.  [also specimen NYS with two specimens on a card].

 

USA: Niagara Falls, G. W. Clinton ("3") several specimens glued to cards (NYS).

 

Canada. Hab. Falls of Niagara. Drummond, Musci Americani (Rocky Mts.) No. 25 (as Gymnostomum recurvirostrum (NY), a tall stemmed group and a small glued to the card, the smaller probably representing the var. minor. Crum and Anderson (1981) indicate that No. 26 of this exsiccat is var. minor. Two other examples seen of No. 25 (both NY) are single collections of one stature. Another specimen from the National Herbarium (CANM) in Ottawa "Falls of Niagara" No. 25. "Welland Co." in pencil.

 

USA: [?] Niagara Falls. legit G. W. Clinton (label of Charles H. Peck, Albany, NY) apparently distributed by Peck to Dr. E. C. Howe (NY). Two collections in fruit representing elongate‑stemmed population and short‑stemmed, both in fruit. Perhaps this particular mode of preserving specimens of Hymenostylium recurvirostrum was to demonstrate the variability of the species, as against naming, for example, short representatives var. inferior, or robust ones with cylindric capsules distinguished by Sullivant and Lesquereux.

 

USA: [?] Niagara Falls, Rev. Francis Wolle, 1873 (NY, in packet of Funaria hygrometrica).

 

[USA?] Niagara Falls, 26 August, 1917, c. fr. A. T. Beals (NY).

 

[USA?] Niagara. c. fr. 1873, Francis Wolle (NY). [ex herb. Eugene A. Rau, Bethlehem, Pa.]

 

[USA?] Niagara Falls. Aug. 21 & 22, 1886. c. fr. Eugene A. Rau, Bethlehem, Pa. (NY)

 

[USA?] Rocks. Niagara. North America. ex Herb. William Mitten (NY).

 

[USA?] On limestone. Niagara. Ruth Schoernhurst, July 17, 1937, c. fr. (NY).

 

CAN: Welland Co., limestone cliffs along Niagara River opposite American Falls, in seepage, Howard Crum no. 26, October 5, 1964 (CANM).

 

A specimen at COLO labeled "Falls Niagara C. W. July 1850", COLO label indicates New York with the possibility of the collector being T. P. James. Specimen mounted on card and hand labeled.

 

As Gymnostmum curvirostrum, "... at Niagara Falls. (Drummond.)," Macoun & Kindberg, 1892 p. 13. 

 

As Gymnostomum curvirostrum Hedw. Queen Victoria Niagara Falls Park.  Cameron 1895. 

 

As Gymnostomum curvirostrum Hedw. Niagara Falls, near the American staircase, Charles Peck in Day, 1883.

 

Podpera (1954) reported an occurrence from "Niagara." Macoun (p. 13, 1892) cited a Drummond specimen from Niagara Falls. Lesquereux & James (1884. pp.54): "Limestone rocks, and on deposits of carbonate of lime or tufa, near springs; very abundant at Niagara Falls."

 

USA:  on rock at base of American Falls, Prospect Point. W. Specht & J. McCandless Oct. 9, 1973 (BUF).

 

CAN: Gorge of the Niagara River, "First Gorge" road into gorge from Queenston, bottom slope, side of gorge, below Lewiston Bridge, wet clay near waterfall R. H. Zander 3723 Sept. 29, 1971 (BUF).

 

CAN: (As Gymnostomum recurvirostrum) Upper Great Gorge of Niagara River, below Table Rock House, near Horseshoe Falls. (spray zone) R. H. Zander Sept. 7, 1971 (BUF); Niagara Falls, near Horseshoe Falls, near bottom of slope, spray zone, on thin soil on rocks with grasses. R. H. Zander 3686b Sept. 7, 1971 (BUF).

 

CAN: Foster's Flats, (Niagara Glen) on the Niagara River, half way down. N-facing, moist, dolomite boulder. RH Zander 63585 Apr 5 1985 (BUF); Niagara Glen, half way up slope from river, Pot Hole Path, boulder, thin soil. R. H. Zander 3732a Sept. 29, 1971 (BUF).

 

USA: Gorge of Niagara R. at Devil's Hole, top of gorge, on limestone.

R Zander 4252a 12 Sept 1975 (BUF). 

 

CAN: Niagara Falls. Forming large tufaceous mass on limestone wall of gorge at the falls. I Herrnstadt & R Zander Mar 2 1977 (BUF).

 

USA.: Ongiara Gorge, between steps and Whirlpool, top of talus slope, base of dolomite caprock in seepage: frozen, with Didymodon tophaceus, northwest facing wall. P. M. Eckel 222386 March 21, l986 (BUF).

 

CAN: Niagara Glen (Foster's Flats). Boulder field. Base of dolomite block, shaded woodland. P.M.Eckel 525086 April 11, 1985 (BUF); Niagara Glen (Foster's Flats) on the Niagara River. Moist dolomite boulder, N-facing, half way up gorge, with Didymodon rigidulus and Desmatodon porteri. With R. Zander. P. M. Eckel 53585 April 5, 1985 (BUF).

 

USA: lower Arch Bridge, upper path USA: Eckel between steps and Whirlpool, seeps, path by caprock, Eckel 321386 (BUF).

 

USA: Niagara River Gorge, Sides Of Sheer Dolomite Cliff, Seepage: Abundant Bryophytes, Upper Old Rr Path Near Top Of Gorge By Schoelkopf Museum. P. M. Eckel 79386 Sept. 21, L985 (BUF).

USA: (as Hymenostylium recurvirostre) just outside spray area of Horseshoe Falls, boulder, thin soil, midslope, with Tortella fragilis, Hyophila involuta, Trichostomum crispulum, Zander 3484d, Oct. 28, 1979 (BUF), on rubble, talus slope, Zander 3491b, Oct. 28, 1979 (BUF); just outside spray area of Horseshoe Falls, vertical rock face of gorge, with Preissia quadrata, Zander 3492a, Oct.28, 1979 (BUF); spray area of Horseshoe Falls, near river, soil under boulder, Zander 3478, Oct.28, 1979 (BUF); spray zone of American Falls, talus, thin soil, crevices of rock, Zander 3444a, Oct. 28, 1979 (BUF); base of island, 1 Nov. 1988, Buck 16354 (BUF, NY).

CAN: Niagara Falls gorge, near the Horseshoe Falls along road to power station. Wet, dripping dolomite walls. R. H. Zander 3651 Aug. 10, 1971 (BUF); Niagara River Gorge, near Horseshoe Falls, down road to powerhouse, Lockport Dolomite, crevices R. H. Zander 3648b Aug. 10, 1971 (BUF).

USA: Luna Island, 1 Nov. 1988, Buck 16331 (NY).

Hyophila involuta (Hook.) Jaeg. & Sauerb. NY CHECK

 

2005 Rare Plant Bryophyte Status List S1

 

[No country] Hab. Niagara Falls, 1873, F. Wolle (NY).

 

USA: Zander, Devil's Hole, lower slopes.

 

USA: Gorge of Niagara River, riverside limestone boulders. R. H. Zander Sept. 12, 1975 (BUF).

 

CAN: Gorge of Niagara River at Niagara Glen, about half way between water and lip of gorge, side of moist, shady limestone boulder. R. H. Zander April 7, 1984 (BUF) .

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. Moist dolomite boulder in solution cavity: the smaller plant is Gymnostomum aeruginosum With R. H. Zander P. M. Eckel 431285 April 5, 1985 (BUF).

 

CAN. gorge of Niagara River, Niagara Glen, near path, floor of woods near river, moist rock surface, R. Zander & P. Eckel, Dec. 4, 2004 (MO).

USA: just outside spray area of Horseshoe Falls, boulder, thin soil, midslope, with Tortella fragilis, Hymenostylium recurvirostrum, Trichostomum crispulum, Zander 3484c Oct. 28, 1979 (BUF), Zander 3480, Oct. 28, 1970 (BUF); spray area of Horseshoe Falls, near river, thin soil in crevice of boulder, Zander 3469, Oct. 28, 1970 (BUF).

Hypnum abietinum Hedw. = Thuidium abietinum (Hedw.) BSG

Hypnum aduncum Hedw. = Drepanocladus aduncus (Hedw.) Warnst.

Hypnum alleghaniense C.Muell. = Thamnobryum alleghaniense (C. Muell.) Nieuwl. 

 

Hypnum brevirostre Ehrh. A Clinton specimen from Devil's Hole (herb no. 41058).

 

Hypnum confervoides Schwaeg. Schwaegr.

 

USA: On rocks, with Radula complanata, second sister Goat island (1873  Clinton, Coll. herb. no. 41072 (BUF).

 

Hypnum cupressiforme Hedw.  NY CHECK

 

USA: Artpark, Eckel 451486 (BUF), (Eckel, l986b).

 

Hypnum curvifolium Hedw. NY CHECK

 

USA: Suspension Bridge (now City of Niagara Falls by the Lower Arch Bridge) archived letter BUF from Charles Peck to George Clinton (collector, from Clinton's  trip May 11, 1867 noted in Clinton's journal)

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: New York, Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring); P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

USA: Erie Co. Grand Island. At delta of creek flowing into Niagara R,

south tip of Grand Island. Ann Glowny 1096 3-28-67(BUF).

 

Hypnum imponens Hedw. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: Navy Island in the Niagara River just S of Niagara Falls, NW end, open gallery, spring-wet Quercus, Ulmus americana, Acer rubrum woods.P. M. Eckel s.n. July 2, 1998(BUF); Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Fallen log; embankment; with Dicranum fuscescens.  Pseudoparaphyllia laciniate. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF); Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. W side; fallen log; with Ptilidium pulcherrimum, Dicranum flagellare. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

 Note this species has costa short and double, and that its stem cortical cells are little or not inflated but especially the few thicker‑walled orange cells at the base of the leaf subtended by a few enlarged thin‑walled hyaline cells (cf. Drepanocladus, which has a costa).

 

Hypnum lindbergii Mitt. (New to NY CHECK)

 

USA: Devil's Hole, upper slopes, Eckel 331585 (BUF, NYS), Eckel, l986a; wet calcareous flat at base of gorge in shrubby river margin, P. M. Eckel Sept. 22, 2004 (BUF); Devil's Hole by the Power Authority, Niagara River Gorge. Upper slopes, calcareous, with Dryopteris marginalis, on rock talus, with

With R. Zander. Dicranella varia. P. M. Eckel 331585 March 16, 1985

(BUF).

 

CAN: Niagara Glen (Foster's Flats) on the Niagara River. On punky log in late snow melt - boulder field. Note there are few rotten log substrates in the flats. With R. Zander P. M. Eckel 531285 April 5, 1985 (BUF).

 

USA: Second of the Three Sisters Islands, east end, on the Niagara River just S of Goat Island. Northern margins by water. P.M. Eckel 881111 Aug. 8, 1988 (BUF); Second Sister, Eckel 86121701 (BUF, Bailey)

 

USA: Erie Co., Grand Island Twp., Buckhorn Island State Park; NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring), P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF).

 

CAN: Dufferin Islands, marshy area on log, Oct. 8, 1989, Eckel  9303175 (BUF).

USA: Second Sister Island, west end, 2 Nov. 1988, Buck 16379 (BUF, NY).

USA: Second Sister Island, east end, thin soil over dolomite flats, open with Salix, Cornus, Betula, Lythrum, Carices, Eckel 86121701, Sept. 12, 1986 (BUF); 2 Nov. 1988, Buck 16404 (BUF, NY).

CAN: Navy Island in the Niagara River just S of Niagara Falls, northwest and northern river margins; "slump meadow" wet red clay eroding embankment. P. M. Eckel s.n. July 15, 1998 (BUF); Navy Island in the Niagara River: 43*03'N,79*01'W, just south of Niagara Falls. Dominant on soil, island west side; Beech-Maple, Carya-Quercus forest; Atrichum undulatum. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF); Navy Island in the Niagara River: 43*03'N, 79*01'W, just south of Niagara Falls. Dominant on logs, interior of island, with Dicranum flagellare; Red Maple forest, wet. P. M. Eckel with R. Zander s.n. Sept. 14, 1998(BUF).

 

Hypnum pallescens (Hedw.) P. Beauv.

USA: Second Sister Island, west end, on trunk of Salix, 2 Nov. 1988, Buck 16388 (BUF, NY).

CAN: Navy Island in the Niagara River 43*03'N, 79*01'W, just south of Niagara Falls. With Hedwigia ciliata on granite glacial erratic; Fraxinus-Carya woods P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Hypnum pygmaeum (BSG) Sull. & Lesq. = Thuidium pygmaeum BSG [note also Hypnum pygmaeum Vill. ex Brid. = Leskeella nervosa (Brid.) Loesk.

 

Hypnum rivulare Brid.(BSG) Bruch =  Brachythecium rivulare  BSG

 

Hypnum rugosum Ehrh. ex Hedw. = Rhytidium rugosum (Hedw.) Kindb.

 

 Hypnum sprucei (Bruch) Spruc. = Amblystegium jungermannioides (Brid.) Giac. fid. Giac.

 

Hypnum subcompactum,  a doubtful name, not in Index Muscorum

 

CAN: Queen Victoria Niagara Falls Park.  Cameron 1895.

 

CAN: "... on rocks and stones below Queenston Heights, Ont. (Macoun.)," Macoun, 1902, p. 312.  CHECK CITATION

 

Hypnum schreberi Willd.

 

[CAN?]: City Niagara Falls. Wood below the Whirlpool. Ex Coll. G. W. Clinton, Buffalo, N.Y. (BUF).

 

A specimen from "wood below the Whirlpool" which Zander has suggested is the wooded moss-rock house area associated with Whirlpool (DeVeaux) steps.

 

Hypnum sullivantii R. Spruce

 

A Clinton specimen from Devil's Hole with fruit (herb no. 38547).

 

Isopterygium elegans (Brid.) Lindb. NY CHECK

 

Kindbergia praelonga (Hedw.) Ochyra

 

As Stokesiella praelonga (Hedw.) Robins.

 

Found by Macoun in 1891 at Foster's Flats, Welland Co. (CANM 183141). Note other records in Cook, 1985. (Crum & Anderson, Ireland etc.)

 

(As Stokesiella praelonga (Hedw.) Robins. in Crum & Anderson, l981.

 

See Crum & Anderson 1981, p. 1077: Canada, Foster's Flats.

 

Leptobryum pyriforme (Hedw.) Wils.

 

CAN: As "Leptobigum pyriforma," Queen Victoria Niagara Falls Park.  Cameron 1895.

 

USA: Luna Island, Niagara Falls [cf. handwriting E. G. Britton, A. A. A. S. meeting] Aug. 21, 1886 (NY, in packet of Funaria hygrometrica). 

 

USA: Buckhorn Island State Park.

 

USA: New York, Erie Co., Grand Island Twp., NW tip Grand Island wet Quercus bicolor, Carya, Juglans nigra, Tilia americana woods just N of marsh. Rotten punky stump, seasonally inundated (spring). P. M. Eckel, Oct. 22, 1997 with R. Zander (BUF), with Pohlia nutans.

 

CAN: Leptobryum vulgare, Queen Victoria Niagara Falls Park.  Cameron 1895.. Doubtful name. Possibly Leucobryum vulgare (= L. glaucum) was meant.

 

Leptodictium humile (P.Beauv.) Ochyra

 

CAN: Ontario, Regnl. Mun.   Niagara, town  of Niagara‑on‑the‑Lake, W side Niagara River on S boundary Fort George. Spring‑inundated soil sub Quercus alba, Quercus rubra forest,  with Brachythecium campestre and Physcomitrium pyriforme. P. M. Eckel, Oct. 4, 1997 BUF.

 

Leptodictium riparium (Hedw.) Warnst.

(As Amblystegium riparium). USA: Second Sister Island, west end, tree roots, wet in high water, Oct. 29, 1988 (BUF).

(As A. riparium) Goat Island in the Niagara River, southeast end, wet soil by riverside. Open, in mats. P. M. Eckel June 3, 1987 (BUF); Dolomite ballast, open eastern end of Goat Island, south side. Barbula unguiculata,Bryum lisae var cuspidatum P.M. Eckel June 4, 1988 (BUF).

 

Leptodictyum trichopodium var. kochii (BSG) Broth.                     

 

Amblystegium kochii, Bruch & Schimp.

 

Leptodictyum trichopodium var. curvipes (BSG) Broth. is reported as excluded from Canada, Ireland et al. mosses of Canada. Macoun (1902) reported the var. curvipes at Niagara Falls, but gave Amblystegium kochii in its synonymy. What was probably found at Niagara was the var. kochii

 

As Amblystegium curvipes BSG "On old logs, Foster's Flats, Niagara Falls, Ont., May 13th, 1901 ... (Macoun.)," Macoun, 1902, p. 302. 

 

Lescuraea patens (Lindb.) H. Arnell & C. Jens.

 

Pseudoleskea atrovirens (Brid.) BSG

 

As Pseudoleskea atrovirens, "On rocks at Niagara Falls (Macoun.)," Macoun & Kindberg, 1892, p. 180. 

 

As Hypnum atrovirens Dicks. and Hypnum filamentosum, Dicks. l.c.

 

Pseudoleskea atrovirens Bruch & Schimp.

 

"On rocks; Niagara Falls and Lake Superior (Macoun)..." (Lesquereux & James, 1884, p. 319), probably in Canada.

 

Leskea gracilescens Hedw.

 

USA: Erie Co. Grand Island. Marshy oak woods on Love Rd., Niagara R.

Tree base in swamp. Ann Glowny 1363 4-20-67 (BUF).

 

Leskea nervosa (Brid.) Myr. = Leskeella nervosa (Brid.) Loeske 

 

Leskea polycarpa Hedw. var. polycarpa NY CHECK

 

Hypnum medium, Drumm. Musc. Bor.‑Am. No. 223.

 

Leskea subobtusifolia C. Muell. & Kindb. in Mac. & Kindb. 1892.

 

USA: RR Bridge, Upper path Eckel 33895 (BUF), Eckel l986.

 

USA: Erie Co., north end of Grand Island, Buckhorn Island State Park, SW portion of Park, we