Preliminary Cryptogamic (Moss, Lichen and Liverwort) Flora of the Canadian and American Gorge at Niagara Falls 1. Mosses

P. M. Eckel
Res Botanica, a Missouri Botanical Garden Web site

http://www.mobot.org/plantscience/ResBot/index.htm

June 21, 2004

 

PRELIMINARY CRYPTOGAMIC (MOSS, LICHEN AND LIVERWORT) FLORA OF THE CANADIAN AND AMERICAN GORGE AT NIAGARA FALLS

1. Mosses

 

P. M. Eckel, Missouri Botanical Garden, PO Box 299, St. Louis, MO 63166-0299 USA

 

 

 

MOSSES

 

Abietinella abietina (Hedw.) Fleisch. = Thuidium abietinum (Hedw.) BSG.

 

Aloina brevirostris (Hook. & Grev.) Kindb.

 

C. Delgadillo showed the region of Niagara Falls on his distribution map, p. 358,  Crum & Anderson 1981.

 

Aloina rigida (Hedw.) Limpr.

 

CAN: As Barbula rigida Hedw, Queen Victoria Niagara Falls Park .Cameron 1895.

 

USA: Zander, Ongiara, lower slopes

 

"Rare and scattered" according to Craig, in Grout's Moss Flora. Niagara, Delgadillo, Bryologist 78:299, 1975. 

 

"Niagara Falls, and two places clearly in Ontario." Crum & Anderson (1981, p. 360). Macoun (1892, p.49) reported a specimen of this species collected "On earth at Niagara Falls, 15th June, 1884 (Macoun)," although Ireland and Cain (1975) did not locate Macoun's specimen on which he based his report.

 

A specimen identified as Barbula rigida from Niagara Falls, Canada by J. Macoun, 1889 (572b), located at Stockholm (S) was redetermined as Aloina brevirostris (H. & G.) Kindb. by both C. Delgadillo (Mar. 1973) and R. Zander (1993) [synoicous, with Barbula convoluta].

 

Amblystegium fluviatile (Hedw.) BSG. NY CHECK

 

CAN: (As Hygroamblystegium fluviatile) Niagara Glen. Ann Glowny 2116 4-13-67 (BUF).

 

USA: Goat Island, Eckel 28686 BUF, NYS. (Eckel, 1986b).

 

 USA: Goat Island on the Niagara River. Dolomite flats just west of the Three Sisters Islands.Wet roots Cornus stolonifera. P. M. Eckel July 5, 1988 (BUF ).

 

USA: (As Hygroamblystegium fluviatile) Three Sisters Islands, wet marly

area on east end of Second Sister.Wm. R. Buck 16395 2 Nov 1988 (BUF), NY).

 Note: these specimens should be compared with Amblystegium tenax.

 Amblystegium jungermannioides (Brid.) Giac. fid. Giac.

= Hypnum sprucei (Bruch) Spruc.

 

USA: (As "A." sprucei Brch.) Goat Island. James in Pr. Am. Phil. Soc. (Peck 1866),

 

USA: (as Hypnum sprucei (Bruch) Spruce)  Goat Is., Niagara Falls, James@, Day 1883. 

 

USA: Goat Island. On the ground. Thomas P. James July, 1850 (BUF).

 

Amblystegium riparium  (Hedw.) BSG = Leptodictium riparium (Hedw.) Warnst.

 

Amblystegium serpens (Hedw.) BSG NY CHECK

 

USA: Suspension Bridge (now City of Niagara Falls by the Lower Arch Bridge) (as Hypnum serpens) "(the fruiting one) also a little of some doubtful Hypnum perhaps laetum or rutabulum and a stem or two of  Mnium": archived letter BUF from Charles Peck to George Clinton (collector, from Clinton's  trip May 11, 1867 noted in Clinton's journal)

 

USA: Eckel, Three Sisters.

 

USA. Goat Island just above Three Sisters Island. Soaking soil by rushing water, low area, calcareous substrate. With R. H. Zander. P. M. Eckel NFlll May 5, 1984 (BUF);  Goat Island,oak-maple forest over limestone; north slope of Goat Island. W. R. Buck 16298 1 Nov. 1988 (BUF, NY).

 

USA: Whirlpool Bridge. At R.R. Bridge under over-pass. Upper path along rim of gorge along dolomite caprock on industrial cinder midden. On cinders.Eckel 231185 3-9-85 (BUF).

 

USA. DeVeaux College woods, hummock, old growth woods, with Fissidens taxifolius, Ceratodon purpureus.  With A. tenax var. spinifolium. P. M. Eckel with R. Zander s.n. May 7, 2000 (BUF).

 

CAN. As Hypnum serpens Hedw., Queen Victoria Niagara Falls Park. Cameron 1895.

 

CAN. Niagara Glen (Foster's Flats). On rock, halfway down to Niagara River, dolomite boulder field. With Tortula mucronifolia, Fissidens bryoides. P.M.Eckel 1374901 April 11, 1985 (BUF).

 

USA: Goat Island, G. W. Clinton, May [NYS]

 

USA: Whirlpool Woods, G. W. Clinton, June [NYS]

 

Amblystegium serpens var. juratzkanum (Schimp.) Rau & Herv.

(In NY CHECK as A. juratzkanum Schimp.)

 

USA: by Robert Moses Pkwy Overpass and Lower Arch Bridge, upper path; moist soil over dolomite. P. M. Eckel with R. H. Zander 191185 June 2, 1985 (BUF).

 

USA: Boulder field, halfway down Whirlpool steps, May 6, 1987, c. fr., Eckel 9302092g (BUF).

 

USA: By Robert Moses Pkwy Overpass and lower Arch Bridge, upper path, moist soil over dolomite, June 2, 1985 Eckel & Zander No. 191185 (BUF)

 

A specimen of Amblystegium minutissimum from Niagara Falls, New York, probably collected by Fr. Wolle, July 18, 1874 (NY), was a particularly small form of this species. An Albany (NYS) specimen of A. T.  Beals' (No. 230, Aug. 26, 1917, as Thuidium pygmaeum "on moist limestone and on small limestone pebbles, Niagara (American) Falls", duplicate from the Wiegmann herbarium at the New York Botanical Garden) is a depauperate, minute form of Amblystegium juratzkanum mixed with Fissidens bryoides. 

USA: base of Goat Island; spray area of Horseshoe Falls, near river, thin soil over rocks, base of falls, with Fissidens cristatus, Zander 3475b, Oct. 28, 1979 (BUF).

USA: Erie Co.  Grand Island. Marshy elm woods at so. end Grand Island. Niagara R. On tree base. Ann Glowny 1296 3-28-67 (BUF).

 

Amblystegium tenax (Hedw.) C.E.O. Jensen

 

= Hygroamblystegium tenax (Hedw.) Jenn. NY CHECK

See Amblystegium fluviatile.

 

USA: (as Hypnum orthocladon P.-Beauv. ) Clinton specimen from Devil's Hole, herb no. 41139.

 

(As Hypnum irriguum Hook. & Wils.) "Niagara Falls ..." (Lesquereux & James, 1884, p. 374).

 

USA: Lewis falls (not Goat) Upper As Amblystedium orthocladon (P. Beauv.) Mac. & Kindb.

 

CAN: "On earth below Queenston Heights, Ont., May 8th, 1901 (Macoun.)," Macoun, 1902, p. 302.

 

USA: (as Hygroamblystegium tenax (Hedw.) Jenn.) Niagara Falls, edge American Falls (not Goat Island). M. Lewis April 12, 1981 (BUF).

 

USA: Goat Island, south shore (Eckel 1990b); weedy bank facing Three Sisters, on Limestone rocks occ. wet, with Impatiens.P. M. Eckel Sept. 9, 1984 (BUF); Dolomite flats just west of the Three Sisters Islands. Wet base Cornus stolonifera. P. M. Eckel June 4, 1988 (BUF); Central Acer-Fraxinus mixed old woods, soil, west end of woods in spray zone of the Horseshoe Falls. Sandy loam. Base of tree.PM Eckel, Marguerite Eckel 9004271 April 15 1990 (BUF).

 

USA: north slope of Goat Island. Wm. R. Buck 16285 1 Nov 1988(BUF, NY).

 

USA: var. tenax. Base of Goat Island at bottom of falls on talus slope. Wm. R. Buck 16356 1 Nov 1988 (BUF, NY).

USA: var. tenax. Goat Island, Terrapin Point, 2 Nov. 1988, Wm. R. Buck, 16440 (NY).

USA: Luna Island ,1 Nov. 1988, Wm. R. Buck 16318 (BUF, NY), 16332 (BUF, NY).

USA. Bath Island (=Green Island) in the Niagara River just N of Goat Island. Generally disturbed area of lawns, base of Ash, river's edge. With Bryum capillare Hedw. P.M. Eckel May 26, 1988 (BUF); Green Island on the Niagara R just N of Goat Island. Old dolomite ballast on S side. Submersed, wet to moist river margin. PM Eckel, Marguerite P. Eckel 9004157 April 15 1990 (BUF).

USA: First Sister Island, western half, 2 Nov. 1988, Wm. R. Buck, 16421 (BUF, NY), 16427 (BUF, NY).

var. tenax. . USA: Second Sister Island, west end, tree roots, wet in high water, Oct. 29, 1988 (BUF); 2 Nov. 1988, Wm. R. Buck, 16382 (NY); var. tenax. wet marly area on east end of Second Sister. Wm. R. Buck 16398 2 Nov 1988 (BUF, NY), 16403 (BUF, NY), 16406 (BUF, NY).

var. tenax. USA: Third Sister Island, west end, 2 Nov. 1988, Wm. R. Buck 16436 (BUF, NY); var. tenax. USA: Third Sister Island, east end, 2 Nov. 1988, Wm. R. Buck 16431 (BUF, NY), Buck 16433 (BUF, NY).

 

USA. Top of Whirlpool Steps, N of the Whirlpool, Niagara Gorge. Moist area. w. R. Zander. P. M. Eckel 880923 Sept. 5, 1988 (BUF).

 

CAN. Gorge of the Niagara River, near Horseshoe Falls, down rd. to power house, on sandstone, near shoreline.R. H. Zander 3658 Aug. 10, 1971 (BUF).

 

Amblystegium tenax var. spinifolium (Schimp.) Crum & Anderson.

 

USA. DeVeaux College woods, hummock, old growth woods, with Fissidens taxifolius, Ceratodon purpureus.  With A. serpens. P. M. Eckel with R. Zander s.n. May 7, 2000 (BUF).

 

CAN. Gorge of the Niagara River, near Horseshoe Falls, near bottom of slope, spray zone, on thin soil on rocks with grasses. R. H. Zander 3682 Sept. 7, 1971 (BUF).

   

Amblystegium trichopodium (Schultz) Hartm.

 

As Leptodictyum trichopodium (Schultz.) Warnst. NY CHECK

USA. (As Leptodictyum trichopodium): Third Sister Island, west end, 2 Nov. 1988, Buck 16435 (NY).

USA. DeVeaux College woods by Whirlpool State Park near gorge of Niagara River. Most frequent moss in damp elevations in bare soil. Old growth, Sugar Maple-Red Oak woods.P. M. Eckel s.n. May 18, 2000 (BUF).

 

CAN. Navy Island, 43*03'N, 79*01'W, just S of Niagara Falls; west side, bark base of tree; Oak woodland. Ceratodon, Dicranum montanum. P. M. Eckel s.n. Sept. 21, 1998 (BUF).

 

Amblystegium varium (Hedw.) Lindb. NY CHECK as var. varium

 

CAN."... quite common on stones and earth at Niagara and Queenston, Ont., 1901 ... (Macoun.)," Macoun, 1902, p. 301‑02.

 

USA: Eckel, RR Bridge, upper slopes.

 

USA: City of Niagara Falls, Whirlpool State Park, in solo sylvatico sterili calcareo, in luco Querci rubri, umbroso, P. M. Eckel, Oct. 4, 1997 (BUF), with Mnium cuspidatum.   

 

CAN: Eckel, Foster's Flats, lower slopes.

 

USA: Eckel 522386, lower slopes, between steps & Whirlpool.

 

USA: Goat Island, Second Sister, side of dolomite boulder, near N end just above waterline, Zander, June 11, 1991 (BUF).

 

USA. Base of Goat Island, spray zone of Horseshoe Falls, near river, soil under boulder, Zander 3478, Oct. 28, 1970 (BUF).

 USA: Goat Island, south side, flats just west of Three Sisters, island margin of Cornus stolonifera (Eckel 1990b).

USA. Gorge of the Niagara River, near the American Falls by Prospect Point, near river, driftwood. R. H. Zander 3744 Sept. 30, 1971 (BUF); thin soil, crevices of limestone boulder. R. H. Zander 3741a Sept. 30, 1971 (BUF).

 

USA: Whirlpool Street at the RR Bridge under the Robt Moses Pkwy overpass, upper path along dolomite caprock. On porous cinder.P.M.Eckel with R. Zander 131685 March 9, 1985 (BUF).

 

USA. Whirlpool Street by the RR Bridge under the overpass, along path along dolomite caprock just below the lip of the gorge. Punky old stump. With R. H. Zander stump.P. M. Eckel 1031185 March 9, 1985 (BUF).

 

USA. Whirlpool State Park, in solo sylvatico sterile calcareo, in luco Querci rubri, umbroso. Mixed with Mnium cuspidatum. P. M. Eckel s.n. Oct. 4, 1997 (BUF).

 

CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Dicranella varia, Fissidens taxifolius, Weissia brachycarpa, Physcomitrium pyriforme, May 27, 1991, Eckel No. 9302250 (BUF).

 

CAN. Niagara River Gorge, near Horseshoe Falls, down road to power house, dry roadside bank, soil. det. P. M. Eckel  1995 R. H. Zander 3655 Aug. 10, 1971 (BUF).

 

CAN. Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Shaded soil; shore; top of embankment; + Barbula unguiculat; Anomod. rugelii.P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF); northwest and northern river margins. + A. serpens, Fiss. taxifol., Didymodon fallax.P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF).

 

Amphidium lapponicum (Hedw.) Schimp.

 

CAN: As  Zygodon lapponicus (Hedw.) BSG "... on damp rocks, Whirlpool Rapids, Niagara Falls, 1901 (Macoun.)," Macoun, 1902, p. 226.

 

Anomodon attenuatus (Hedw.) Hueb. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park .Cameron 1895.

 

CAN: Niagara Glen (Foster's Flats) Just N Of The Whirlpool, Base Of Dolomite Boulder, N-Facing Aspect, With Myurella siberica, Lophocolea heterophylla. P. M. Eckel 321186 April 11, L985 (BUF).

 

CAN: Navy Island in the Niagara River, northeast side, dry fringe meadow; soil, characteristic of vegetation of meadow between crest and shrubby forest border. P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF).

 

USA: Zander, Ongiara, lower Zander 3521a (NYS, BUF) (Eckel, l986).

 

USA: talus, between steps and Whirlpool, Eckel 221386 BUF.

USA: First Sister Island, western half, Eckel 880723, July 8, 1987 (BUF); 2 Nov. 1988, Buck 16429 (BUF, NY).

Anomodon minor (Hedw.) Fuernr. NY CHECK

 

Anomodon obtusifolius Bruch & Schimp.

 

CAN: (As Anomodon obtusifolius),  Queen Victoria Niagara Falls Park.Cameron 1895.

 

CAN: (As Anomodon obtusifolius) "Trunks of trees in Upper Canada (Ontario) and about Niagara Falls. (Drummond.)," Macoun & Kindberg, 1892, p. 171. 

 

CAN: Hab. Trunks of trees in Upper Canada; and about the Falls of Niagara. Drummond, Musci Americani no. 163 (as A. viticulosum) (NY). 

 

USA: Ongiara Gorge of Niagara River, bottom of W facing slope, between Devil's Hole and the Whirlpool, thin soil on boulder, wet area. R. H. Zander April l7, 1971 (BUF).

 

Anomodon rostratus (Hedw.) Schimp. NY CHECK,

 

CAN: Queen Victoria Niagara Falls Park.Cameron 1895.

 

USA: New York, on rock, vic. Lewiston, Clifford A. LaMere No. 405, April 21, 1962 (NYS).

 

CAN: Niagara Glen (Foster's Flats). Boulder field. Base of dolomite block, shaded woodland. P.M.Eckel 725086 April 11, 1985 (BUF).

 

CAN: Niagara Glen, just N of Whirlpool. Vertical face of limestone boulder, mesic Acer, Tsuga canadensis woods, bottomland. P.M.Eckel with R. Zander NF9 Feb. 19, 1983 (BUF).

 

USA: Whirlpool St. by the RR Bridge, upper path along the dolomite caprock..dolomite surface. With Amblystegium varium & Orthotrichum anomalum. With R. Zander P. M. Eckel 131785 March 9, 1985 (BUF).

 

USA: Ongiara Gorge of Niagara River, bottom of W facing slope, between Devil's Hole and Whirlpool, thin soil on boulder, wet area. R. H. Zander April 17, 1971 (BUF).

USA: First Sister Island, western half, Eckel 880726, July 8, 1987 (BUF); 1 Nov. 1988, Buck 16335 (BUF, NY).

USA: Third Sister Island, northeast end, on boulder, Eckel, Sept. 29, 1988 (BUF).

Anomodon rugelii (C.M.) Keissl. NY CHECK

 

CAN: As Anomodon apiculatus BSG ex Sull. in Gray following Crum & Anderson p. 870. Queen Victoria Niagara Falls Park. Cameron 1895 (CAN).

 

CAN: Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Shaded soil; shore; top of embankment; + Amblyst. varium; Thuidium delicatulum.

P. M. Eckel with R. Zander s.n. May 13, 1998

 

Anomodon viticulosus (Hedw.) Hook. & Tayl. NY CHECK

 

From a letter from Leo Lesquereux to George Clinton Oct. 16, 1865 (BUF archives):

    "About Anomodon viticulosus, you will find it in plenty against a large

rock near the base of that footpath descending from Goat Island to the top

of the American falls. You know that there are two ways of going there: the

broad way by stairs and aside of it, on the right a small slippery foot

path on the slope of the Island; it is on this foot path about halfway down

that said Anomodon grows. I never found it in America except at this place.

James collected it also there, &c"

 

USA: "Rocks. Goat Island, Lesqx. Various localities about Niagara Falls, G. W. Clinton." (Peck 1866).

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895. 

 

CAN: Niagara Falls; Whirlpool, Niagara River, Canadian side.  Day 1883

 

USA: Rocks & trees, Niagara Falls, July 18, 1874 (NY) Herb. Coe Finch Austin (probably all colls. 1874 are Wolle's). 

 

Hab. ad rupes humidas Niagarae cataractae. No. 359 [Sullivant & Lesquereux, Musci Bor.‑Amer., ed. 2, 359] (NY)

 

USA: Ad rupes humidas Niagarae cataractae. Musci Bor. Am. Sull. & Lesq. Ed. 2. 1865. Sull. & Lesq. (BUF).

 

USA: Niagara Falls. G.W.Clinton (BUF).

 

Hab. ad rupes humidas Niagarae cataractae. No. 359 [Sullivant & Lesquereux, Musci Bor.‑Amer., ed. 2, 359] (NYS)

 

USA: Hab. Niagara Falls [N.Y. penciled in later], 1873, leg. F. Wolle (NY).

 

CAN: On limestone rocks, Niagara River. J. Macoun, May 18, 1901 (NY).

 

USA: Devil's Hole, limestone on rocks in dry creek valley, R. & I. Duell, May 19, 1981 (NY).

 

USA: boulder field, half way down Whirlpool steps, with Bryum lisae var. cuspidatum, Campylium chrysophyllum, Tortella tortuosa, Plagiopus oederiana, Marchantia polymorpha, Mnium thomsonii, Encalypta procera, Gymnostomum aeruginosum, May 6, 1987, Eckel No. 9302092a (BUF).

 

CAN: Eckel, Foster's Flats and boulder‑field, half way down.

 

CAN: Foster's Flats (Niagara Glen). on the Niagara River. Abundant, over dolomite blocks. Boulder field, medis hemlock-thuja bottomland.

With R. Zander. P. M. Eckel 431285 April 5, 1985 (BUF).

 

Clinton specimen, " Niagara Falls" herb no. 36141

 

"Shaded rocks; Niagara Falls, sterile ..." (Lesquereux & James, 1884 p. 306). Note Clinton's journal may indicate that Lesquereux himself collected the sterile specimen. 

 

USA: On stone Niagara Falls. "(This is a Eur. specimen. I had no time to look for those of Niagara)" [L. Lesqu.]

Herbarium number: 36142  (BUF).

 

"Shaded rocks; Niagara Falls, sterile. (Lesq. & James)," Macoun & Kindberg, 1892, p. 171. 

 

USA: Crest of the gorge of the Niagara River, just N of Whirlpool State Park. With Barbula unguiculata w. R. H. Zander. P. M. Eckel 880919 Sept. 5, 1988 (BUF).

 

Astomum muhlenbergianum (Sw.) Grout.

 

USA: Eckel slope sewage treatment plant, Upper Gt.Gorge. Eckel 17686 (BUF), (Eckel, l986b).

 

Atrichum angustatum Bruch & Schimp.

 

Catharinea angustata Brid.

 

CAN: On earth, Queenston, Ont. May 13, 1901. John Macoun, coll, det. by Dr. N. C. Kindberg. Canadian Mosses No. 181 (US, distributed by the National Herbarium of Canada).

 

CAN: "... on earth in woods, Queenston Heights, Niagara River, Ont., 1901.  (Macoun.)," Macoun, 1902, p. 262. 

 

CAN: Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Dry ground, low elevation in mucky ground.P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).

 

Atrichum oerstedianum (C. Muell.) Crum.

 

USA: Artpark, Eckel & Eckel 914486 (BUF, NYS)  (Eckel, l986b).

 

ONT:(as A. undulatum var. oerstedianum (C.M.) Crum): Navy Island in the Niagara River: 43*03'N,79*01'W, just south of Niagara Falls.Dominant on soil, island west side; Beech-Maple, Carya-Quercus forest; Hypnum lindbergii. P. M. Eckel with R. Zander s.n. Sept. 14, 1998 (BUF).

 

Atrichum undulatum (Hedw.) P.-Beauv.

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.

 

CAN: (as var. altecristatum Ren. & Card.) Navy Island in the Niagara River just S of Niagara Falls, central wet Quercus, Ulmus americana, Acer rubrum woods, with Fissidens taxifolius, Eurhynchium hians.P. M. Eckel s.n. July 15, 1998

 

USA: (as var. altecristatum Ren. & Card.) DeVeaux Woods State Park, deeply shaded virgin woods Quercus rubra, Acer saccharum, Quercus alba, little herbaceous growth. Near building complex. Soil. Sparse. Eckel, June 1, 2006 (MO).

 

Aulacomnium heterostichum (Hedw.) BSG

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895

 

Aulacomnium palustre (Hedw.) Schwaegr.

 

CAN: Queen Victoria Niagara Falls Park.Cameron 1895. see Disc. sub Sphagnum. 

 

"... in wet woods, Niagara Falls, Ont., 1901 (Macoun.)," Macoun, 1902, p. 261.

 

Erie Co.: Grand Island. Wet soil in thicket, Buffalo Ornithological Refuge, Love Rd., Grand Isl., Niagara River. Ann Glowny 10502 7-5-72 (BUF).

 

Barbula convoluta Hedw. NY CHECK

 

CAN: Queen Victoria Niagara Falls Park. Cameron 1895.

 

CAN: "On earth at Queenston Heights, Ont., May 8th, 1901. (Macoun.)," Macoun, 1902, p. 213. 

 

A specimen at NY Barbula convoluta was "on earth, Queenston Heights" collected by J. Macoun, 8/5 1901. The notation 15 in ink in the upper left corner and in pencil: "=13".

 

CAN: [[A specimen identified as Barbula rigida from Niagara Falls, Canada by J. Macoun, 1889 (572b), located at Stockholm (S) was redetermined as Aloina brevirostris (H. & G.) Kindb. by both C.  Delgadillo (Mar. 1973) and R. Zander (1993) [synoicous, with Barbula convoluta] ‑ see A. rigida above. 

 

Macoun goes on to say: "All eastern references found under this species in Cat. Can. Musci, 57, belong here. All western to the next species" [= B.  chysopoda, C. Muell. & Kindb. was described earlier in Macoun, 1892, p.  57], which is a synonym of B. convoluta.

 

USA: Whirlpool Pk, upper slopes,  Zander May 15, l976 (BUF, NYS) Eckel, l986.

 

USA: Eckel 1622386 between steps & Whirlpool, seep in caprock

 

CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Dicranella varia, Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Physcomitrium pyriforme, May 27, 1991, Eckel No. 9302250 G (BUF). 

 

USA: Green Island on the Niagara River, Just N of Goat Island. Old Dolomite ballast on S side, wet to moist river margin. With Pottia t.,Barbula u.,Phascum c.,Bryum spp. PM Eckel, Marguerite P. Eckel 9004159 April 15 1990 (BUF).

 

USA: Erie Co.:  Grand Island. In crevices of boulder by marsh, southern tip of Grand Island, Niagara River. Ann Glowny 1787 3-27-68 (BUF).

 

Barbula fallax Hedw. NY CHECK

USA: spray zone of American Falls, talus, midslope, pathside, thin soil, Zander 3451, Oct. 28, 1979 (BUF).

USA: Devil's Hole Gorge of Niagara River, below Devil's Hole, bottom of W facing slope, top of rock wall, thin soil. R.H. Zander April 17, 1971 (BUF); Lewiston. Devil's Hole by the power authority on the Niagara River, upper slopes, calcareous ss, shales, dolomite on talus-cobble, open, full shade half day from surrounding  cliffs. P.M.Eckel with R. Zander 132285 March 16, 1985 (BUF).

 

USA: (As Didymodon fallax var. fallax) Second Sister Island, west end, tree roots, wet in high water, Oct. 29 1988 (BUF); Second of the Three Sisters Islands in the Niagara River just S of Goat Island. Muck alluvium, west end, with Didymodon tophaceus, humectated tree roots. P.M. Eckel 881101 Oct. 29, 1988 (BUF).

 

ONT: (As Didymodon fallax var. fallax) Navy Island in the Niagara River, just south of Niagara Falls, northwest and northern river margins. Dry fringe meadow, soil between crest and shrubs, with Fiss. taxifolius. P. M. Eckel with R. Zander s.n. July 15, 1998 (BUF).

 

Barbula reflexa (Brid.) Brid. NY CHECK

 

CAN: As Barbula recurvifolia Mitt, Queen Victoria Niagara Falls Park.Cameron 1895.

 

As B. recurvifolia Schimp.; Lesq. & James, Mosses of N. America, 122. No. 4 [64?] Geological and Natural History Survey of Canada no. (201). Niagara Falls. (Austin). [hand written:] dry rocks. N. Falls, July 1874 (NY). = Didymodon ferrugineus (Besch.) Hill det. R. H. Zander, 2001: probably Canada.

 

Macoun (p. 54, 1892) cited a specimen collected by Coe Finch Austin from Niagara Falls, Ontario.

 

As Barbula recurvifolia, Schimp, "Niagara Falls (Austin)..." (Lesquereux & James, 1884, p.122), no country cited.

 

Barbula unguiculata Hedw. NY CHECK

 

Barbula apiculata, Hedw.

 

Barbula lanceolata, Hedw.

 

Barbula stricta Hedw.

 

Trichostomum indigens Ren. & Card.

 

[USA?] Niagara Falls. Rev. Francis Wolle (NY).

 

CAN: Queen Victoria Niagara Falls Park.Cameron 1895.

 

USA [?] Whirlpool, Niagara River, Day 1883.

 

USA: Zander, Eddy Basin, upper slopes.

 

USA: Niagara River Gorge, Sides Of sheer Dolomite Cliff, Seepage: Abundant Bryophytes, Upper Old Rr Path Near Top Of Gorge By Schoelkopf Museum. Det. Zander. P.M.Eckel 19286 Sept. 21, 1985 (BUF).

 

USA: City of Niagara Falls, Whirlpool St. at Lower Arch (R.R.) Bridge, upper path just below rim of gorge, dolomite caprock, vertical rock wall, thin soil in moist fine cracks. R. Zander & P. M. Eckel 8612211 Dec. 12, 1986 (BUF). USA: Zander, Ongiara Gorge, lower slopes 

 

USA: between steps & Whirlpool Eckel 521386 BUF.

 

USA: Eckel 226386, Devil's Hole, above.

USA: Spray area of American Falls, talus, midslope soil, trailside, Zander 3454a, Oct. 28, 1979 (BUF).

CAN: Base of wooded embankment overlooking the Horseshoe Falls.  In spray on calcareous clay in lawn, with Dicranella varia, Fissidens taxifolius, Amblystegium varium, Weissia brachycarpa, Physcomitrium pyriforme, May 27, 1991, Eckel No. 9302250A (BUF). 

 

CAN: Dufferin Islands, mats on moist soil, April 4, 1993, c. fr., Eckel 9304223 (BUF).

 

CAN: Navy Island in the Niagara River, just south of Niagara Falls, western side of island. Shaded soil; shore; top of embankment; + Amblyst. varium; Thuidium delicatulum. P. M. Eckel with R. Zander s.n. May 13, 1998 (BUF).<