BFNA Title: Homaliadelphus
Author: I. Sastre de Jesús 
Date: August 24, 2008
Edit Level: R 
Version: 1

Bryophyte Flora of North America, Provisional Publication
Missouri Botanical Garden

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XX. HOMALIADELPHUS Dixon & P. de la Varde, Rev. Bryol. Lichenol. 2. 4: 142. 1932. * [Genus Homalia and Greek adelphos, brother, alluding to a similarity with Homalia]


Plants small, flat, glossy. Stem creeping, sparsely branched to irregularly pinnate; flagelliform branches absent. Paraphyllia present. Stem and branch leaves wide-spreading, smooth, flat, slightly asymmetric, apex rounded, small lobe at adaxial base of lateral leaves; margins entire or slightly denticulate; costa absent or short-double.  Inner perichaetial leaves oblong from a sheathing base [ovate at base turning subulate], obtuse, apex entire to crenulate proximally [smooth], serrulate; apical cells quadrate to rhombic, basal cells rectangular. Sexual condition dioicous (male plants epiphytic on female plants). [Seta 4--6 mm. Capsule oblong-cylindric; exostome teeth lanceolate, smooth to slightly striate at base; endostome with low basal membrane. Spores 15--20 \um]


Species 2 (1 in the flora): North America, Mexico, Asia (China, India, Japan).


The characteristic adaxial basal leaf lobe of Homaliadelphus easily separates this genus from other members of the Neckeraceae. In North America this genus is restricted to calcareous rocks, although in India it occurs as an epiphyte.


1. Homaliadelphus sharpii (R. S. Williams) Sharp, Castanea 9: 107. 1944


Homalia sharpii R. S. Williams, Bryologist 34: 20, plate 2. 1931


Plants less than 3 cm. Stem and branch leaves obovate to nearly rounded, 0.5--1.5(--2) mm; apical cells rounded-quadrate to short-oblong, 6--9 x 6 \um, distal medial cells irregularly rhombic, 10 x 10 \um, basal cells 20--27 x 9 \um, basal marginal cells short-rhomboidal to fusiform-elongate, slightly pitted, ca. 9 x 8 \um.


Shaded calcareous rocks, cliffs or soil, hardwood-pine forests; 200--500 m; Ala., Mo., Tenn., Va.; Mexico; Asia (China, India, Japan).