Michael Grayum 9578 Det. by: A. R. Smith, 1992 Phenology: 15 December 1989 08:58N 83:18W 90 m PUNTARENAS, BUENOS AIRES Valle del Térraba. Locally common in pockets of soil on spray-soaked limestone wall of waterfall at roadside.
Michael Grayum 9175 Det. by: R. Moran, 1989 Phenology: 9 December 1988 08:58N 83:28W 40 m PUNTARENAS, OSA Valle del Térraba. Locally abundant on weedy banks in mango plantation on hillside. Stems reddened.
Michael Grayum 8077 Det. by: R. Moran, 1988 Phenology: 27 February 1987 09:48N 83:58W 1700 m CARTAGO, EL GUARCO Valle del Reventazón. Erect plants, locally abundant on damp, weedy slopes. Stems red-tinged, articulate.