Michael Grayum 10589 Det. by: A. R. Smith 1995 Phenology: 7 December 1993 08:49N 83:11W 900-1000 m PUNTARENAS, OSA Fila Costeña. Growing on steep rock bank of creek in primary forest; rhizome erect.
J.F. Morales 1959 Det. by: J. F. Morales, V 1994 Phenology: 6 November 1993 09:50N 84:06W 2150 m SAN JOSE, ASERRI Z.P. Cerros de Escazú Cerros Escazú-La Carpintera. Terrestre. Soros verde crema.
Michael Grayum 8905 Det. by: R. Moran, 1989 Phenology: 9 September 1988 10:00N 83:24W 180 m LIMON, MATINA Z.P. Barbilla Cordillera de Talamanca. Epilithic or terrestrial in primary forest on rocky ridge. Abundant here.
Gerardo Herrera 2194 Det. by: R. Moran 1992 Phenology: 15 October 1988 10:00N 83:25W 200 m LIMON, MATINA Z.P. Barbilla Cordillera de Talamanca. Terrestre. Abundante. Follaje suave, verde-tenue. Soros café.
James Solomon 19118 Det. by: R. Moran, 1992 Phenology: 31 August 1990 10:57N 85:29W 600 m GUANACASTE, LA CRUZ P.N. Guanacaste Cordillera de Guanacaste. Forest understory.
II INBio 103 Det. by: R. Moran, 1991 Phenology: 2 July 1989 10:57N 85:29W 600 m GUANACASTE, LA CRUZ P.N. Guanacaste Cordillera de Guanacaste. Helecho terrestre.
II INBio 108 Det. by: R. Moran, 1991 Phenology: 2 July 1989 10:57N 85:29W 600 m GUANACASTE, LA CRUZ P.N. Guanacaste Cordillera de Guanacaste. Helecho terrestre.
Manuel Lobo 54 Det. by: A. Rojas, V 1996 Phenology: 11 February 1995 10:57N 85:29W 800 m GUANACASTE, LA CRUZ P.N. Guanacaste Cordillera de Guanacaste. Helecho terrestre. Soros cafés.