Michael Grayum 9281 Det. by: M. Grayum, II 1989 Phenology: 24 January 1989 08:47N 82:58W 1140 m PUNTARENAS, COTO BRUS J.B. Rob. & Cat. Wilson Valle de Coto Colorado. Uncommon terrestrial plants in forest on ridge. Petioles and rachis glossy black.
Michael Grayum 10639 Det. by: M. Grayum, VIII 1994 Phenology: 9 December 1993 08:49N 83:11W 800-950 m PUNTARENAS, OSA Fila Costeña. Terrestrial and epilithic in primary forest on limestone ridge. Stipe and rachis blackish, glossy. Fronds radicant at apex.
Abelardo Chacón 393 Det. by: M. Grayum, IX 1990 Phenology: 23 September 1989 09:17N 83:13W 1700 m PUNTARENAS, BUENOS AIRES R.I. Ujarrás Cordillera de Talamanca. Helecho de 0.25-0.35 m. Tallos café morados (vivípara). Soros café.
Jens Bittner 1728 Det. by: A. Rojas, X 1995 Phenology: 11 March 1993 09:17N 83:16W 1530 m PUNTARENAS, BUENOS AIRES R.I. Ujarrás Cordillera de Talamanca. Herb to 0.2 m.
Gerardo Herrera 3491 Det. by: M. Grayum, I 1990 Phenology: 19 September 1989 09:17N 83:14W 1400 m PUNTARENAS, BUENOS AIRES R.I. Ujarrás Cordillera de Talamanca. Terrestre, esporangios café.
Alexande Rojas 2570 Det. by: A. Rojas, V 1996 Phenology: 25 April 1996 09:41N 84:23W 290-350 m SAN JOSE, PURISCAL Z.P. La Cangreja Cerros de Puriscal. Terrestre con luz media y materia orgánica escasa.
Gerardo Rivera 389 Det. by: R. Moran, 1992 Phenology: 12 June 1990 10:07N 84:08W 2599 m HEREDIA, BARVA P.N. Braulio Carrillo Cordillera Central. Helecho terrestre. Esporas ferrugíneas.
Darin Penneys 697 Det. by: A. Rojas, XI 1995 Phenology: 21 August 1995 10:43N 85:06W 600-700 m ALAJUELA, UPALA Cordillera de Guanacaste. Growing on boulders. Plantlets rooting from frond apex.