Michael Grayum 9253 Det. by: M. Grayum, II 1989 Phenology: 12 December 1988 08:39N 83:12W 80 m PUNTARENAS, GOLFITO R.N.V.S. Golfito Valle de Coto Colorado. Epilithic on large rocks in cascading creek in primary forest. Petioles fleshy, deep blackish-green toward base, not glossy.
Michael Grayum 9151 Det. by: M. Grayum, II 1989 Phenology: 9 December 1988 08:59N 83:28W 780 m PUNTARENAS, OSA Fila Costeña. Epilithic on rocky slopes and dripping rock faces along small forest creek.
Jens Bittner 1270 Det. by: R. Moran, V 1992 Phenology: 25 February 1992 09:26N 83:03W 310 m LIMON, TALAMANCA P.N. Cordillera de Talamanca Cordillera de Talamanca. Epihythe on tree in 2 m.
Michael Grayum 9446 Det. by: M. Grayum, V 1989 Phenology: 8 May 1989 10:26N 84:01W 50 m HEREDIA, SARAPIQUI Estación Biológica La Selva Llanura de San Carlos. Terrestrial ferns, the rhizome short- creeping. Small population on steep, muddy E bank of Quebrada Arboleda, just upstream from Quebrada El Surá.