Michael Grayum 9152 Det. by: R. Moran, 1990 Phenology: 9 December 1988 08:58N 83:28W 300 m PUNTARENAS, OSA Valle del Térraba. Terrestrial on steep, rocky slope in primary forest. Uncommon here. Probably conspecific with MG 9159.
Sary Rojas 41 Det. by: M. Grayum, VII 1994 Phenology: 16 September 1992 08:59N 83:16W 100 m PUNTARENAS, BUENOS AIRES R.I. Boruca Valle del Térraba. Helecho terrestre. Creciendo a la orilla de una quebrada.
Michael Grayum 9868 Det. by: M. Grayum, VII 1990 Phenology: 9 July 1990 08:59N 83:28W 300-400 m PUNTARENAS, OSA Valle del Térraba. Terrestrial in rocky soil in disturbed primary forest. Rhizome short-creeping. Petiole and rachis dark brown, semi- glossy.