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J.F. Morales 4259 Det. by: P. Sánchez, I 1996 Phenology: LR 2 June 1995 09:43N 84:10W 1600-1800 m SAN JOSE, ACOSTA Fila Bustamante. Arbol de 7 m x 22 cm DAP. Botones florales marrón. Frutos verde-amarillo.
Patrick Harmon 349 Det. by: P. Sánchez, I 1996 Phenology: FL 9 July 1994 10:25N 84:45W 600 m ALAJUELA, SAN RAMON Cordillera de Tilarán. Subcanopy tree 18 m x 25 cm dbh. Cylindrical trunk with no buttresses; tannish, smooth bark. Opposite leaves thick and leathery, underside brownish. Buds and spent flowers clustered in leaf axils.