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1. This research was supported by National Geographic Society Grant 4816-92. We gratefully acknowledge the work of B.-E. van Wyk, Rand Afrikaans University, Johannesburg, who provided the analyses of sugar nectars. We also thank Peter Bernhardt, Dee Paterson-Jones, and Kim Steiner for helpful comments during the preparation of this paper.

2. Compton Herbarium, National Botanical Institute, Private Bag X7, Claremont 7735, South Africa.

3. B. A. Krukoff Curator of African Botany, Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166-0229, U.S.A.


Abstract | Materials and Methods | Results | Discussion | Literature