Research Staff

Discovering a new fern species of Polystichum (Dryopteridaceae) endemic to a single karst cave in northern Guangxi, China.

Li-Bing Zhang, Ph.D.
Curator
Co-Director, Flora of China Project
Herbarium

office phone: + 1 (314) 577-9454
fax: +1 (314) 577-9596
email: libing.zhang@mobot.org;
gonggashan@gmail.com

Missouri Botanical Garden
4344 Shaw Blvd.
St. Louis, MO 63110
USA

Education
Ph.D., Johannes Gutenberg University at Mainz, Germany, 2002
M.S., Chengdu Institute of Biology, the Chinese Academy of Sciences, 1990
B.S., Lanzhou University, China, 1987

Management of the Flora of China Phylogeny, systematics, and evolution of ferns, lycophytes, and flowering plants Biogeography of intercontinental disjunctions of plants Systematics and evolution of high mountain plants and cave plants

Available upon request

Research Emphases Systematics of pteridophytes, especially Athyriaceae, Dryopteridaceae (e.g., Arachniodes, Ctenitis, Cyrtomium, Dryopteris, Pleocnema, Polystichum), Polypodiaceae, Pteridaceae, Pteridryaceae, Tectariaceae, and lycophytes Lycopodiaceae and Selaginellaceae Systematics, phylogeny, and biogeography of selected seed plant groups, e.g., Cucurbitales (e.g, Anisophyllaceae), Fabaceae (e.g., Campylotropis, Lespedeza, Desmodieae, Sophora), Oleaceae (e.g., Jasminum), Primulaceae (e.g., Primula, Soldanella), Rosaceae (e.g, Rosa)
New families/subfamilies/genera described

New families Didymochlaenaceae Ching ex Li Bing Zhang & Liang Zhang, 2015 Pteridryaceae Li Bing Zhang, X.M. Zhou, Liang Zhang & N.T. Lu, 2018
New subfamilies Lycopodiastroideae Li Bing Zhang & X.M. Zhou (Lycopodiaceae), 2022 Boreoselaginelloideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Gymnogynoideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Lycopodioidoideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Pulvinielloideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Selaginelloideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Selaginoidoideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Sinoselaginelloideae Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023
New genera of ferns & lycophytes (chronological order) Draconopteris Li Bing Zhang & Liang Zhang (Pteridryaceae), 2016 Malaifilix Li Bing Zhang & Schuettp. (Pteridryaceae), 2016 Gastoniella Li Bing Zhang & Liang Zhang (Pteridaceae), 2017 Hovenkampia Li Bing Zhang & X.M. Zhou (Polypodiaceae), 2017 ×Woodsimatium Li Bing Zhang, N.T.Lu & X.F.Gao (Woodsiaceae), 2020 Shahashia Li Bing Zhang & Liang Zhang (Ophioglossaceae), 2020 Ellipinema Li Bing Zhang & Liang Zhang (Polypodiaceae), 2020 ×Ellipisorus Li Bing Zhang & Liang Zhang (Polypodiaceae), 2020 Brownseya Li Bing Zhang, L. Sheph., D.K.Chen, X.M. Zhou & H.He (Lycopodiaceae), 2022 Goswamia Li Bing Zhang & Liang Zhang (Ophioglossaceae), 2022 Haukia Li Bing Zhang & Liang Zhang (Ophioglossaceae), 2022 Whittieria Li Bing Zhang & Liang Zhang (Ophioglossaceae), 2022 ×Apseudathyrium X.Wan & Li Bing Zhang (Athyriaceae), 2022 ×Osmunimunda X.Wan & Li Bing Zhang (Osmundaceae), 2022 ×Glaphyrocyclosorus X.Wan & Li Bing Zhang (Thelypteridaceae), 2022 Boonkerdia Li Bing Zhang, Pollawatn, X.M. Zhou & Liang Zhang (Polypodiaceae), 2023 Calligrammitis Parris, Sundue, Li Bing Zhang, X.M. Zhou & Ralf Knapp (Polypodiaceae), 2023 Devolia Li Bing Zhang, X.M. Zhou, Jian Jun Yang & Ralf Knapp (Polypodiaceae), 2023 Glabrigrammitis Li Bing Zhang, X.M. Zhou, Jian Jun Yang & Parris (Polypodiaceae), 2023 Grammitastrum (E.Fourn.) Parris, Sundue, Li Bing Zhang & X.M. Zhou (Polypodiaceae), 2023 Howeogrammitis Parris, Sundue, Li Bing Zhang & X.M. Zhou (Polypodiaceae), 2023 Nanogrammitis Parris, Li Bing Zhang, X.M. Zhou & Liang Zhang (Polypodiaceae), 2023 Rouhania Li Bing Zhang, X.M. Zhou, Jian Jun Yang & Liang Zhang (Polypodiaceae), 2023 Thalassogrammitis Parris, Sundue, Li Bing Zhang & X.M. Zhou (Polypodiaceae), 2023 Afroselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Austroselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Boreoselaginella (Warb.) Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Chuselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Ericetorum (Jermy) Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Korallia Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Kungiselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Lepidoselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Megaloselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Pulviniella (Li Bing Zhang & X.M. Zhou) Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Sinoselaginella Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023 Valdespinoa Li Bing Zhang & X.M. Zhou (Selaginellaceae), 2023
Selected Publications

I. Books
Zhang L-B. 2021. View From the Forest and Caves: Travels of a Botanist in Asia. St. Louis: Missouri Botanical Garden Press. Beijing: China Agricultural Science & Technology Press. 245 pp. Zhang L-B. 2016. [Chinese Translation of] McNeill J, Barrie, F.R., Buck, W.R., Demoulin, V., Greuter, W., Hawksworth, D.L., Herendeen, P.S., Knapp, S., Marhold, K., Prado, J., Prud'homme van Reine, W.F., Smith, G.F., Wiersema, J.H. & Turland, N.J. (eds.). 2012. International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011. Bratislava: International Association of Plant Taxonomy. Zhang L-B. 2013. Flora of China Illustrations volume 2-3 (Lycophytes & Ferns), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2013. Flora of China Illustrations volume 20-21 (Asteraceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2012. Flora of China Illustrations volume 19 (Cucurbitaceae through Valerianaceae with Annonaceae and Berberidaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2012. Flora of China Illustrations volume 23 (Acoraceae through Cyperaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2011. Flora of China Illustrations volume 10 (Fabaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2010. Flora of China Illustrations volume 25 (Orchidaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2009. Flora of China Illustrations volume 7 (Menispermaceae through Capparaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2009. Flora of China Illustrations volume 11 (Oxalidaceae through Aceraceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2008. Flora of China Illustrations volume 13 (Clusiaceae through Araliaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2008. Flora of China Illustrations volume 12 (Hippocastanaceae through Theaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2007. Flora of China Illustrations volume 20 (Poaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2007. [Chinese Translation of] McNeill J, FR Barrie, HM Burdet, V Demoulin, DL Hawksworth, K Marhold, DH Nicolson, J Prado, PC Silva, JE Skog, JH Wiersema, and NJ Turland (eds.). 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Guang-Hua Zhu, L-B Zhang. 2006. Flora of China Illustrations volume 14 (Apiaceae through Ericaceae), edited by Wu Zhengyi, Peter H. Raven & Hong Deyuan. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2002. Phylogeny, biogeography and systematics of Soldanella L. and Primula L. sect. Auricula Duby (Primulaceae) based on molecular and morphological evidence. Goettingen: Cuvillier Verlag. 157 pp.
II. Book chapters
Zhang L-B. 2016. Eucommiaceae. Pp. 193–196 In: Kubitzki K (ed.), The Families and Genera of Vascular Plants. Berlin: Springer-Verlag. Euasterids, edited by Kadereit JW. Zhang L-B, Wu S-G, Xiang J-Y, Xing F-W, He H, Wang F-G, Lu S-G, Dong S-Y, Barrington D, Iwatsuki K, Christenhusz MJM, Mickel JT, Kato M, Gilbert MG. 2013. Dryopteridaceae. In Wu Z-Y, Raven PH, Hong D-Y eds. Flora of China, vol. 2-3. Beijing: Science Press, St. Louis: Missouri Botanical Garden Press, 542–724. Zhang L-B & DS Barrington. 2013. Polystichum. Pp. 629–713 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B & DS Barrington. 2013. Cyrtomium. Pp. 561–n: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B & K Iwatsuki. 2013. Lycopodiaceae. Pp. 13–36 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B & WC Taylor. 2013. Isoëtaceae. Pp. 35–36 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B & NJ Turland. 2013. Equisetaceae. Pp. 67–72 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B & G Yatskievych. 2013. Psilotaceae. P. 81 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B, Wu SG, Xiang JY, Xing FW, He H, Wang FG, Lu SG, Dong SY, Barrington D, Iwatsuki K, Christenhusz MJM, Mickel JT, Kato M, Gilbert MG. 2013. Dryopteridaceae. Wu ZY, Raven PH, Hong DY eds. Flora of China, vol. 2–3. Beijing: Science Press, St. Louis: Missouri Botanical Garden Press, 542–724. He H, L-B Zhang & D Barrington. 2013. Dryopteris sect. Acrorumohra (H. Itô) Li Bing Zhang & H. He. Pp. 613–615 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Wu S-G, J-Y Xiang, L-B Zhang & D Barrington. 2013. Dryopteris sect. Nothoperanema Li Bing Zhang. Pp. 618–620 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Wang F-G, F-W Xing, L-B Zhang & D Barrington. 2013. Dryopteris sect. Acrophorus (C. Presl) Li Bing Zhang. Pp. 620–623 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Wang F-G, F-W Xing, L-B Zhang & D Barrington. 2013. Dryopteris sect. Diacalpe (Blume) Li Bing Zhang. Pp. 623–626 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Wang F-G, F-W Xing, L-B Zhang & D Barrington. 2013. Dryopteris sect. Peranema (D. Don) Li Bing Zhang. Pp. 626–627 in: Wu Z-Y, PH Raven & D-Y Hong (eds.). Flora of China. vol. 2–3. Beijing: Science Press and St. Louis: Missouri Botanical Garden Press. Zhang L-B. 2004. Huperziaceae. Pp. 1–54. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6(3), edited by Zhang X-C. Zhang L-B. 2004. Lycopodiaceae. Pp. 55–85. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6(3), edited by Zhang X-C. Zhang L-B. 2004. Isoëtaceae. Pp. 220–223. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6(3), edited by Zhang X-C. Zhang L-B. 2004. Equisetaceae. Pp. 224–243. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6(3), edited by Zhang X-C. Zhang L-B. 2004. Psilotaceae. Pp. 244–245. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6(3), edited by Zhang X-C. Zhang L-B. 2001. Polystichum sect. Metapolystichum. Pp. 52–60. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 5 (2), edited by Kung H-S. Zhang L-B. 2001. Polystichum sect. Neopolystichum. Pp. 84–117. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 5 (2), edited by Kung H-S. Kung H.-S and L-B Zhang. 2001. Polystichum sect. Lasiopolystichum. Pp. 60–85. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 5 (2), edited by Kung H-S.
III. Systematics, phylogeny, and biogeography of ferns and lycophytes
Lu TN, L Zhang, NT Thanh, LC Toan, L-B Zhang. 2023. Three new species and one new record of the fern genus Polystichum (Dryopteridaceae) from Vietnam. Phytotaxa 607(1): 085–094. Fang S-L, Y-X Li, L-B Zhang, Z-R He, X-M Zhou. 2023. Selaginella subchaetoloma (Sect. Heterostachys; Selaginellaceae), a new species from Southwest China. Phytotaxa 603 (3): 224–234. Wan X, R Knapp, D-K Chen, H He, , L-B Zhang. 2023. Taxonomic notes on some East Asian pteridophytes. Phytotaxa 597(2): 128–140. Zhou X-M, J-J Yang, R Pollawatn, R Knapp, B Parris, M Sundue, TA Ranker, Z-L Liang, L Zhou, NT Lu, TT Luong, X Wan, Z-R He, J Zhao, L Zhang, L-B Zhang. 2023. A global phylogeny of grammitid ferns (Polypodiaceae) and its systematic implications. Taxon 73: xx-xx. DOI: https://doi.org/10.1002/tax.12992. Yang J-J, B Parris, R Knapp, L-B Zhang, X-M Zhou. 2023. Circumscription of the grammitid fern genus Oreogrammitis (Polypodiaceae) with the description of three new genera, Calligrammitis, Devolia, and Glabrigrammitis. Phytotaxa 597(1): 28–36. Wang B, Q-W Sun, Z-H Wang, L-B Zhang. 2023. Polystichum kuankuoshuiense (sect. Sphaenopolystichum, Dryopteridaceae), a new fern from Guizhou, China. Phytotaxa 592 (2): 144–150. Xu K-W, Y-X Liu, Y-F Chen, L-B Zhang. 2023. Hymenasplenium hovenkampii (Aspleniaceae), a new bipinnatisect fern species from Lesser Sunda Islands of Indonesia. Phytotaxa 595(1): 120–124. Wan X, L Zhang, S Lehtonen, H Tuomisto, D-W Zhang, X-F Gao, L-B Zhang. 2023. Five long-distance dispersals shaped the major intercontinental disjunctions in Tectariaceae s.l. (Polypodiales, Polypodiopsida). Molecular Phylogenetics and Evolution 186: 107845. Wan X, X-M Zhou, L Zhang, Z-R He, L-B Zhang. 2023. Proposal to conserve the name Selaginella (Selaginellaceae) with a conserved type. Taxon 72 (2): 429–430. Shang H, Z-Q Xue, Z-L Liang, M Kessler, R Pollawatn, TN Lu, Y-F Gu, X-P Fan, Y-H Tan, L Zhang, X-M Zhou, X Wan, L-B Zhang. 2023. Splitting one species into 22: an unusual tripling of molecular, morphological, and geographical differentiation in the fern family Didymochlaenaceae (Polypodiales). Cladistics 39: 273–292. DOI: https://doi.org/10.1111/cla.12539. Shang H, L-B Zhang. 2023. Systematics of the fern genus Didymochlaena (Didymochlaenaceae). Systematic Botany 48(1): 110–139. Lu TN, H-J Wei, LD Vuong, LC Toan, L-B Zhang, L Zhang. 2023. Cyrtomium calcis sp. nov. and six new records of the shield fern family (Dryopteridaceae) from Vietnam. Phytotaxa 583(1): 015–026. Wang B, Q-W Sun, Z-H Wang, L-B Zhang. 2022. Polystichum kuankuoshuiense (sect. Sphaenopolystichum, Dryopteridaceae), a new fern from Guizhou, China. Phytotaxa 592(2): 144–150. Wan X, L-B Zhang. 2022. Three new fern names of hybrid genera and four new combinations in Athyriaceae, Osmundaceae, and Thelypteridaceae. Phytotaxa 575(1): 104–108. Wan X, L Zhang, L-B Zhang. 2022. Validation and lectotypification of the fern combination Whittieria engelmannii (Ophioglossaceae). Phytotaxa 567 (2): 205–206. Wan X, L-B Zhang. 2022. Global new taxa of vascular plants published in 2021 Biodiversity Science 30 (8): 22116, pages 1–8. Nie L-Y, L Zhang, Z-L Liang, R. Pollawatn, Y-H Yan, NT Lu, R Knapp, X Wan, D Cicuzza, X-X Chen, H-F Chen, A-H Wang, Y-J Liao, F-G Wang, L-B Zhang. 2022. Phylogeny, character evolution, and biogeography of the fern genus Bolbitis (Dryopteridaceae). Molecular Phylogenetics and Evolution 178 (2023): 107633. DOI: https://doi.org/10.1016/j.ympev.2022.107633. Wang B, Sun Q-W, Wang Z-H, Chen C-L, Xu W-F, L-B Zhang. 2022. Polystichum suiyangense (sect. Sphaenopolystichum, Dryopteridaceae), a new fern from Guizhou, China. Phytotaxa 567 (1): 093–099. He H, L-B Zhang. 2022. Polystichum gonggashanense (Dryopteridaceae): a new fern from Sichuan, China. Phytotaxa 559 (2): 207–212. Liang Y-Y, L-B Zhang. 2022. Polystichum neocavernicola (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Phytotaxa 559 (1): 095–098. Rajapaksha R, L-B Zhang, DKNG Pushpakumara, DSA Wijesundara. 2022. An analysis of the current status and future prospects of Sri Lankan pteridophytes towards a new dimension. Biologia. DOI: https://doi.org/10.1007/s11756-022-01139-y. Zhang L, L-B Zhang. 2022. Phylogeny, character evolution, and classification of the fern family Ophioglossaceae based on Sanger sequence data, plastomes, and morphology. Molecular Phylogenetics and Evolution 173 (2022): 107512. Zhou X-M, J Zhao, J-J Yang, T Le Pechon, L Zhang, L-B Zhang. 2022. Plastome structure, evolution, and phylogeny of Selaginella. Molecular Phylogenetics and Evolution 169 (2022) 107410. Liang Z-L, Z-Y Yu, Y-M Shui, W-H Chen, B Tunckol, L Zhang, L-B Zhang. 2022. Polystichum jinpingense (subg. Haplopolystichum; Dryopteridaceae), a new fern from southeastern Yunnan, China. Phytotaxa 539 (1): 069–073. Qiu Y-L, L Zhang, L-B Zhang, X-M Zhou, L Zhang. 2022. Athyrium aberrans (Athyriaceae), a new species of the lady ferns from southeastern Xizang, China, based on morphological and molecular evidence. Phytotaxa 533 (3): 165–172. Petchsri S, T Jaruwattanaphan, L-B Zhang. 2022. A new Ophioglossum species from Thailand. Phytotaxa 533 (3): 158–164. Liang Z-L, L Zhang, X-F Gao, L-B Zhang. 2021 [2022]. Chromosome study of 38 fern species in Dryopteridaceae, Hypodematiaceae, Polypodiaceae, and Pteridaceae from China. Indian Fern Journal 38: 198–218. Zhang L-B, X-M Zhou. 2021 [2022]. A new classification of Lycopodiales based on molecular phylogenetics, morphology, and spore ornamentation. Indian Fern Journal 38: 125–136. Zhang L-B. 2021. Fern & Lycophytes. Poster. Missouri Botanical Garden Press. Xu, K-W, TN Lu, X-M Zhou, L Zhang & L-B Zhang. 2021. Asplenium quangbinhense sp. nov. and three new records of Asplenium (Aspleniaceae) from Vietnam. Phytotaxa 521 (2): 113–120. Chen D-K, X-M Zhou, CJ Rothfels, L Shepherd, R Kapp, L Zhang, NT Lu, X-P Fan, X-F Gao, H He & L-B Zhang. 2021. A global phylogeny of Lycopodiaceae (Lycopodiales) with the description of a new genus, Brownseyia, from Oceania. Taxon https://doi.org/10.1002/tax.12597. Duan, Y-F, M Li, K-W Xu, L Zhang & L-B Zhang. 2021. Protect China's cave habitats. Science 374 (6568), https://doi.org/10.1126/science.abm5389. Wan, X & L-B Zhang. 2021. Global new species of vascular plants published in 2020 Biodiversity Science> 29 (8): 1003–1010. Fan, X-P, TN Lu, L Zhang & L-B Zhang. 2021. Hypodematium chrysolepis comb. nov. from Africa and H. eglandulosum sp. nov. from Asia (Hypodematiaceae). Phytotaxa 524 (2): 099–106. Du, X-Y, J-M Lu, L-B Zhang, J Wen, L-Y Kuo, CM Mynssen, H Schneider & D-Z Li. 2021. Simultaneous diversification of Polypodiales and angiosperms in the Mesozoic. Cladistics 37: 518–539. He, M, Z-R He, L-B Zhang & X-M Zhou, 2021. Selaginella qingchengshanensis (sect. Heterostachys; Selaginellaceae), a new species from Sichuan, China. Phytotaxa 522 (4): 285–293. Fan, X-P, TN Lu, C-X Li, R Knapp, H He, X-M Zhou, L Zhang, X-F Gao & L-B Zhang. 2021. Phylogeny, biogeography, and character evolution in the fern family Hypodematiaceae. Molecular Phylogenetics and Evolution. https://doi.org/10.1016/j.ympev.2021.107340 Xu K-W, L-L Wang & L-B Zhang. 2021. Taxonomic revision of the Asplenium wrightii complex (Aspleniaceae) with reinstatement of A. alatulum and A. subcrenatum. Phytokeys 172: 75–91. Xu, K-W, TN Lu, X-M Zhou, L Zhang & L-B Zhang. 2021. Asplenium quangbinhense sp. nov. and three new records of Asplenium (Aspleniaceae) from Vietnam. Phytotaxa 521 (2): 113–120. Shang H, Z-L Liang & L-B Zhang. 2021. Taxonomy of the fern genus Didymochlaena (Didymochlaenaceae) from Asia and Pacific islands based on morphological and molecular evidence with the description of four new species and one new status. Phytotaxa 479(1): 071–082. Fan X-P, L Zhang, R Knapp, H He, X-F Gao & L-B Zhang. 2021. Hypodematium shingii sp. nov. (Hypodematiaceae; Polypodiales): Replacing the misapplied “H. crenatum”, a widespread fern from Asia. Phytotaxa Fan X-P, L-B Zhang & L Zhang. 2021. Selliguea wusugongii (Polypodiaceae), a new fern species from southeastern Xizang, China based on morphological and molecular evidence. Phytotaxa 480(1): 057–068. Liang Z-L, Z-R He & LL-B ZhangL. 2021. Polystichum dongchuanense and P. menglaense (subg. Haplopolystichum; Dryopteridaceae), two new ferns from Yunnan, China. Phytotaxa 479(1): 114–120. Zhang L & L-B Zhang. 2020. ×Ellipisorus nom. nov. for ×Lepinema (Polypodiaceae) and ×Ellipisorus inaequibasis com. nov. from China. Phytotaxa 464(4): 299–300. Zhang L & L-B Zhang. 2020. Fern nothogenera and ×Lepinema, a new hybrid genus between Ellipinema and Lepisorus (Polypodiaceae). Phytotaxa 455(4): 262–266. Shang H, Z-Q Xue, Y-F Gu & L-B Zhang. 2020. Revision of the fern genus Didymochlaena (Didymochlaenaceae) from Madagascar. Phytotaxa 459(4): 252–264. Fan X-P, X-F Gao & L-B Zhang. 2020. Taxonomy of the fern genus Hypodematium (Pteridophyta: Hypodematiaceae) from Japan. Phytotaxa 455(2): 161–166. Liang Z-L, Y-M Shui, W-H Chen, Z-Y Yu, L Zhang & L-B Zhang. 2020. Leptochilus gracilis (Polypodiaceae), a new fern species from southeastern Yunnan, China based on morphological and molecular evidence. Phytotaxa 468(1): 112–120. Zhang L & L-B Zhang. 2020. Fern nothogenera and ×Lepinema, a new hybrid genus between Ellipinema and Lepisorus (Polypodiaceae). Phytotaxa 455(4): 262–266. Camara-Leret et al. 2020. New Guinea has the world’s richest island flora. Nature https://doi.org/10.1038/s41586-020-2549-5. Ye Z-Y, NT Lu, L Zhang, X-M Zhou, L-B Zhang. 2020. Selaginella coriaceifolia (sect. Heterostachys; Selaginellaceae), a new lycophyte species from central Vietnam. Phytotaxa 453(2): 121–129. Tunckol B, H. 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New combinations in the fern genus Leptochilus (Polypodiaceae). Phytotaxa 374: 172–176. Xu K-W, L Zhang, NT Lu & L-B Zhang. 2018. Asplenium serratifolium (Aspleniaceae), a new fern species from central Vietnam based on morphological and molecular evidence. American Fern Journal 108: 65–75. Han M-Q, Y Liu & L-B Zhang. 2018. Eight new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from limestone caves in Guangdong and Yunnan, China, with reference to species diversity in the karst terrains at high elevations in subtropical areas. Phytotaxa 365: 145–168. Gang G-D & Zhang L-B. 2018. Tectaria moranii (Tectariaceae), a new fern Costa Rica. Phytotaxa 357(3): 230–234 Xu K-W, Zhang L., Lu TN, Zhou X-M, He H, Luong TT, Knapp R, Liao W-B & Zhang L-B. 2018. Nine new species of Hymenasplenium (Aspleniaceae) from Asia. Phytotaxa 358: 1–25. Xu K-W, Zhou X-M, Liao W-B & Zhang L-B. Hymenasplenium perriei (Aspleniaceae), a new fern species from Fiji. 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Zhou, X.-M., Zhang, L., Lu, N.T., Gao, X.-F., L-B Zhang. 2018. Pteridryaceae: A new fern family of Polypodiineae (Polypodiales) including taxonomic treatments. Journal of Systematics and Evolution 56: 148–173. Xu K-W, Zhou X-M, Zhang L-B & Liao W-B. 2018. Hymenasplenium hastifolium sp. nov. (Aspleniaceae) from a karst cave in western Guangxi, China. Phytotaxa 333: 281–286. Zhang, L & L-B Zhang. 2018. Phylogeny and systematics of the brake fern genus Pteris (Pteridaceae) based on molecular (plastid and nuclear) and morphological evidence. Molecular Phylogenetics and Evolution 118: 265–285. Duan, Y-F, M. Kropf & L-B Zhang. 2017. Polystichum zhijinense (subg. Haplopolystichum; Dryopteridaceae), a new cave species of Polystichum from Guizhou, China. Phytotaxa 331: 124–130. Zhang, L., T.N. Lu, X.-M. Zhou & L-B Zhang. 2017. Three new species of the fern genus Pteridrys (Tectariaceae) from Vietnam. Phytotaxa 324: 279–287. Zhou X-M & L-B Zhang. 2017. Nuclear and plastid phylogenies suggest ancient intersubgeneric hybridization in the fern genus Pyrrosia (Polypodiaceae), with a classification of Pyrrosia based on molecular and non-molecular evidence. Taxon 66: 1065–1084. Lu, TN, Y-F Duan & L-B Zhang. 2017. Polystichum alluvium (subg. Haplopolystichum; Dryopteridaceae), a new cave species from Guizhou, China, with reference to new distribution records of P. leveillei. Phytotaxa 323: 088–092. Duan, Y-F, NT Lu, L Zhang & L-B Zhang. 2017. Polystichum luteoviride (subg. Haplopolystichum; Dryopteridaceae), a new, highly endangered cave species from Guizhou, China. Phytotaxa 313: 296–300. Zhang, L-B. 2017. On the terms in the International Code of Nomenclature for algae, fungi and plants. Journal of Sichuan Normal University 40(5): 1–8. Chen, D-K, Zhou X-M, H He & L-B Zhang. 2017. Spinulum lioui, a new species referred as to Lycopodium neopungens (Lycopodiopsida: Lycopodiaceae) in China. Phytotaxa 307: 161–163. Zhang, L., Zhou, X.-M., Chen, D.-K., Schuettpelz, E., Knapp, R., Lu, N.T., Luong, T.T., Dang, M.T., Duan, Y.-F., He, H., Gao, X.-F., L-B Zhang. 2017. A global phylogeny of the fern genus Tectaria (Tectariaceae: Polypodiales) based on plastid and nuclear markers identifies major evolutionary lineages and suggests repeated evolution of free venation from anastomosing venation. Molecular Phylogenetics and Evolution 114: 295–333. Zhou, X.-M., Zhang, L., Chen, C.W. Li, C.-X., Huang, Y.M., Chen, D.-K., Lu, N.T., Cicuzza, D., Knapp, R., Luong, T.T., Nitta, J.H., Gao, X.-F., L-B Zhang. 2017. A plastid phylogeny and character evolution of the Old World fern genus Pyrrosia (Polypodiaceae) with the description of a new genus: Hovenkampia (Polypodiaceae). Molecular Phylogenetics and Evolution 114: 271–294. Hennequin, S., G. Rouhan, A. Salino, Y.-F. Duan, M.-C. Lepeigneux, M. Guillou, S. Ansell, T. Almeida, L.-B. Zhang & H. Schneider. 2017. Global phylogeny and biogeography of the fern genus Ctenitis (C.Chr.) C.Chr. (Dryopteridaceae), with a focus on the Indian Ocean region. Molecular Phylogenetics and Evolution 112: 277–289. Tang, G.-D., Huang, L., Li, J.-Y., He, Z.-L. & Zhang, L.-B. 2017. Polystichum hastipinnum (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangdong, China. Phytotaxa 309: 66–72. Zhou X-M, C-W Chen & L-B Zhang. 2017. Pyrrosia annamensis comb. nov. (Polypodiaceae) from Southeast Asia and lectotypification of Cyclophorus rhomboidalis (Polypodiaceae). Phytotaxa 309: 090–094 Zhang, L.-B. 2017. From excellence to eminence: Relaunching Annals of the Missouri Botanical Garden. Annals of the Missouri Botanical Garden 102: 1–2. Zhang, L.-B. 2017. The PPG I classification and pteridophytes of China. Biodiversity Science 25: 340–342. He, H. & L.-B. Zhang. 2017. Polystichum tiandengense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guangxi, China. Novon 25: 153–157. Zhou X-M & L-B Zhang. 2017. Dendrolycopodium verticale comb. nov. (Lycopodiopsida: Lycopodiaceae) from China. Phytotaxa 295: 199–200. Xu X-G, X-M Zhou & L-B Zhang. 2017. Huperzia wusugongii nom. nov. (Lycopodiopsida: Huperziaceae) from West China. Phytotaxa 296: 098–099. Peng Y-L, X-F Gao, L-B Zhang. 2017. Youngia jiulongensis (Asteraceae), a new species from Sichuan, China. Novon 25: 298–301. Miao X-Y, C-X Li & L-B Zhang. 2017. Polystichum duyunense (subg. Haplopolystichum; Dryopteridaceae), a new cave fern from Guizhou, China. Phytotaxa 292: 296–300. Duan, Y-F, S Hennequin, G Rouhan, B Bassuner, and L-B Zhang. 2017. A taxonomic revision of the fern genus Ctenitis (Dryopteridaceae) from Africa and the Western Indian Ocean. Annals of the Missouri Botanical Garden 102: 3–86. Zhang L, X-M Zhou, NT Lu & L-B Zhang. 2017. Phylogeny of the fern subfamily Pteridoideae (Pteridaceae; Pteridophyta), with the description of a new genus: Gastoniella. Molecular Phylogenetics and Evolution 109: 59–72. Xu X-G & L-B Zhang. 2016. Pleocnemia siamensis (Dryopteridaceae), a new fern species from southern Thailand. Phytotaxa 289: 88–92. PPG (L-B Zhang is one of the 26 organizers) I. (2016) A community-derived classification for extant lycopods and ferns. Journal of Systematics and Evolution 54: 563–603. Pongkai P, L-B Zhang & R Pollawatn. 2016. Hypodematium boonkerdii sp. nov. (Hypodematiaceae; Pteridophyta) from Thailand. Phytotaxa 286: 193–197. Chen, D-K, H He & L-B Zhang. Request for a binding decision on whether Huperzia rubricaulis (Alderw.) Holub and H. rubicaulis S.K. Wu & X. Cheng (Lycopodiopsida: Huperziaceae) are sufficiently alike to be confused. Taxon 65: 1184–1185. Peng, Y-L & L-B Zhang. 2016. Humbertacalia madagascarensis sp. nov. (Asteraceae: Senecioneae) from Madagascar. Phytotaxa 283: 291–294. Lu TN, L Zhang, D.T. Nguyen & L-B Zhang. 2016. Polystichum quangbinhense (subg. Haplopolystichum; Dryopteridaceae), the southernmost cave species of Polystichum from central Vietnam. Phytotaxa 283: 295–299. Li C-X, NR Patel & L-B Zhang. Polystichum clarinervium (subg. Haplopolystichum; Dryopteridaceae), a new fern from Emei shan, China. Phytotaxa 280: 271–277. Han M-Q, Y Liu & L-B Zhang. Seven new species of Polystichum (subg. Haplopolystichum; Dryopteridaceae) from southern China. Phytotaxa 280: 201–221. Kadereit, JW, DC Albach, F Ehrendorfer, Mè Galbany-Casals, N Garcia-Jacas, B Gehrke, G Kadereit, N Kilian, JT Klein, MA Koch, M Kropf, C Oberprieler, MD Pirie, CM Ritz, M Röser, K Spalik, A Susanna, M Weigend, E Welk, K Wesche, L-B Zhang & MS Dillenberger. 2016. Which changes are needed to render all genera of the German flora monophyletic? Willdenowia 46: 39–91. Zhang, L., Schuettpelz, E., Rothfels, C., Zhou, X.-M., Gao, X.-F., L-B Zhang. 2016. Circumscription and phylogeny of the fern family Tectariaceae based on plastid and nuclear markers, with the description of two new genera: Draconopteris and Malaifilix (Tectariaceae). Taxon 65: 723–738. Le Péchon T, H He, L Zhang, X-M Zhou, X-F Gao, L-B Zhang. 2016. Using a multilocus phylogeny to test morphology-based classifications of one of the largest fern genera Polystichum (Dryopteridaceae). BMC Evolutionary Biology 16:55. Le Péchon, T, L Zhang, X-M Zhou, H He, X-F Gao, L-B Zhang. 2016. A well-sampled phylogenetic analysis of the polystichoid ferns (Dryopteridaceae) suggests a complex biogeographical history involving both boreotropical migrations and recent transoceanic dispersals. Molecular Phylogenetics and Evolution 98: 324–336. Zhang L, NT Lu, and L-B Zhang. 2015. Pteris langsonensis (Pteridaceae), a new brake fern species from Lang Son Province, northern Vietnam. Phytotaxa 238: 283–287. Zhou, X-M, L-J Jiang, L Zhang, X-J He, X-F Gao, Z-R He, and L-B Zhang. 2015. Spore morphology of Selaginella (Selaginellaceae) from China and its systematic significance. Phytotaxa 237: 1–67. Zhou, X-M and L-B.Zhang. 2015. A classification of Selaginella (Selaginellaceae) based on molecular (chloroplast and nuclear), macromorphological, and spore features. Taxon 64: 1117–1140. Zhang L, NT Lu, and L-B Zhang. 2015. Leptochilus oblongus (Polypodiaceae), a new fern species from northern Vietnam. Phytotaxa 234: 195–198. Zhou X-M, Z-R He, L. Zhang, and L-B Zhang. 2015. Selaginella chuweimingii (Selaginellaceae) sp. nov. from Yunnan, China. Phytotaxa 231: 283–288. Zhou, X-M, CJ Rothfels, L Zhang, Z-R He, T Le Péchon, H He, NT Lu, R Knapp, D Lorence, X-J He, X-F Gao, L-B Zhang. 2015. A large-scale phylogeny of the lycophyte genus Selaginella (Selaginellaceae: Lycopodiopsida) based on plastid and nuclear loci. Cladistics. DOI: 10.1111/cla.12136. Duan Y-F and L-B Zhang. 2015. Notes on the fern flora of Fiji: Synonymization of Ctenitis minima and Dryopteris waiwaiensis with Tectaria dissecta (Tectariaceae) and Deparia boryana (Athyriaceae), respectively. Phytotaxa 218: 197–199. Sun Q-W, Zhao J-H & L-B Zhang. 2015. Selaginella daozhenensis (Selaginellaceae), a new lycophyte from a limestone cave in northern Guizhou, China. Phytotaxa 207: 187–192. Duan Y-F and L-B Zhang. 2015. Validation of Ctenitis jinfoshanensis (Dryopteridaceae) and Lepisorus simulans (Polypodiaceae) for fern flora of China. Phytotaxa 205: 299–300. Sessa E, L-B Zhang, H Väre, and A Juslén. 2015. What we do (and don’t) know about ferns: Dryopteris (Dryopteridaceae) as a case study. Systematic Botany 40: 387–399. Zhang L, CJ Rothfels, A Ebihara, E Schuettpelz, T Le Péchon, P Kamau, H He, X-M Zhou, J Prado, A Field, G Yatskievych, X-F Gao, L-B Zhang. 2014. A global plastid phylogeny of the brake fern genus Pteris (Pteridaceae) and related genera in the Pteridoideae. Cladistics 31: 406–423. Lu NT, L Zhang, and L-B Zhang. 2014. Polystichum hagiangense (subg. Haplopolystichum; Dryopteridaceae), a new fern species from Ha Giang Province, northern Vietnam. Phytotaxa 175: 293–297. Zhang L, X-M Zhou, X-F Gao, L-B Zhang. 2014. Pteris barklyae (Pteridaceae): A new combination and neotypification of Adiantum pallens for the fern flora of the Indian Ocean Islands. Phytotaxa 174: 116–118. Zhang L-B, Q-W Sun, and H He. 2014. Selaginella wangpeishanii (Selaginellaceae), a new lycophyte from a limestone cave in Guizhou, China. Phytotaxa 164: 195–199. Zhang L-B. and H-M Liu. 2014. Dryopteris damingshanensis (Dryopteridaceae): new fern in subgenus Nothoperanema from Guangxi, China. Novon 23: 119–122. Lu TN, L Zhang, and L-B Zhang. 2014. Ten new records of Polystichum (Dryopteridaceae) for Vietnam and lectotypification of Polystichum atroviridissimum and P. fimbriatum. Phytotaxa 164: 115¬–123. Zhang L-B. and H He. 2013 [2014]. Book review of Ferns and Fern Allies of Taiwan by R. Knapp. American Fern Journal 103: 191–192. Duan Y-F and L-B Zhang. 2014. Nomenclatural notes on Tectaria blepharorachis (comb. nov.) and T. fibrillosa (Tectariaceae; Pteridophyta) for Malagasy fern flora. Phytotaxa 162: 107–110. Duan Y-F and L-B Zhang. 2014. Megalastrum oppositum and Ctenitis canacae (Dryopteridaceae): A new combination and a new synonym, respectively, for Mascarene fern flora. Phytotaxa 159 (4): 295–297. Zhang L, X-F Gao and L-B Zhang. 2013. Polystichum hainanicola, sp. nov. (sect. Haplopolystichum; Dryopteridaceae) from Hainan, China. Phytotaxa 85: 9–14. Zhang L, X-F Gao and L-B Zhang. 2013. Polystichum hubeiense (sect. Haplopolystichum; Dryopteridaceae), a new fern species from Hubei, China. Annales Botanici Fennici 50: 107–110. Zhang L-B and L. Zhang. 2012. The inclusion of Acrophorus, Diacalpe, Nothoperanema, and Peranema in Dryopteris (Dryopteridaceae): The molecular phylogeny, systematics, and nomenclature of Dryopteris subgenus Nothoperanama. Taxon 61: 1199-1216. Zhang L-B, L Zhang, S-Y Dong, EB Sessa, X-F Gao, and A Ebihara. 2012. Molecular circumscription and major evolutionary lineages of the fern genus Dryopteris (Dryopteridaceae). BMC Evolutionary Biology 12:180. Zhang L-B. 2012. Reducing the fern genus Dryopsis to Dryopteris and the systematics and nomenclature of Dryopteris subgenus Erythrovariae section Dryopsis (Dryopteridaceae). Phytotaxa 71: 17-27. Zhang L-B. 2012. Taxonomic and nomenclatural notes on the fern genus Polystichum (Dryopteridaceae) from China. Phytotaxa 60: 57-60. Luo Q and L-B Zhang. 2012. Dryopteris jiucaipingensis (Dryopteridaceae), a new spcies in D. sect. Hirtipedes from Guizhou, China. Novon 22: 183-185. Zhang L-B, P-S Wang and X-Y Wang. 2012. Dryopteris liboensis (Dryopteridaceae), a new species in D. sect. Erythrovariae from Guizhou, China. Novon 22: 256-259. Zhang L-B, P-S Wang and X-Y Wang. 2012. Selaginella longistrobilina (Selaginellaceae), a new species from Guizhou, China and Selaginella prostrata, a new combination and its lectotypification. Novon 22: 260-263. Luo Q and L-B Zhang. 2012. Woodsia guizhouensis (Woodsiaceae), a new species from a limestone area in Guizhou, China. Novon 22: 191-195. He H, Y-Q Yang and L-B Zhang. 2012. Polystichum loratum (Dryopteridaceae), a new fern from Guizhou, China. Novon 22: 166-169. He H and L-B Zhang. 2012. Polystichum oblanceolatum, a new species in sect. Haplopolystichum (Dryopteridaceae) from Guangxi, China. Novon 22: 160-165. Zhang L, L-B Zhang and J Liu. 2012. Polystichum yaanense (Dryopteridaceae), a remarkable new species from Sichuan, China. Novon 22: 244-249. Luo Q and L-B Zhang. 2012. Polystichum tiankengicola (Dryopteridaceae), a new species from a karst sinkhole from Guizhou, China. Novon 22: 186-190. Zhang L-B and P-S Wang. 2012. Polystichum normale (Dryopteridaceae), a new species in section Crucifilix from China. Novon 22: 250-255. Zhang L-B and H He. 2012. Polystichum perpusillum, sp. nov. (sect. Haplopolystichum, Dryopteridaceae), a new fern species from Guizhou, China. Annales Botanici Fennici 49(1-2): 67-74. Zhang L-B and H He. 2011. Polystichum fengshanense, sp. nov. (sect. Haplopolystichum, Dryopteridaceae) from karst caves in Guangxi, China based on morphological, palynological, and molecular Evidence. Systematic Botany 36: 854-861. He H and L-B Zhang. 2011. Polystichum cavernicola, sp. nov. (sect. Haplopolystichum, Dryopteridaceae) from a karst cave in Guizhou, China and its phylogenetic affinities. Botanical Studies 52: 121-127. He H and L-B Zhang. 2010. Pteris xiaoyingae, sp. nov. (sect. Pteris) from a karst cave in China based on morphological and palynological evidence. Systematic Botany 35: 695-700. He H and L-B Zhang. 2010. Polystichum kungianum, sp. nov. (sect. Mastigopteris, Dryopteridaceae) from Chongqing, China. Botanical Studies 51: 395-401. Zhang L-B, H He, and Q Luo. 2010. Polystichum puteicola, sp. nov. (sect. Haplopolystichum, Dryopteridaceae) from a karst sinkhole in Guizhou, China based on molecular, palynological, and morphological evidence. Botanical Studies 51: 127-136. Zhang L-B and H He. 2010. Polystichum speluncicola sp. nov. (sect. Haplopolystichum, Dryopteridaceae) based on morphological, palynological, and molecular evidence with reference to the non-monophyly of Cyrtogonellum. Systematic Botany 34: 13-19. Zhang L-B and H He. 2009a. Polystichum minutissimum, sp. nov. (sect. Haplopolystichum, Dryopteridaceae): The smallest Polystichum found in a karst cave in China. Botanical Studies 50: 353-358. Zhang L-B and H He. 2009b. Polystichum weimingii sp. nov. (sect. Metapolystichum, Dryopteridaceae) from southern Yunnan, China. Systematic Botany 34: 13-16. Zhang L-B and H He. 2009c. Polystichum peishanii (sect. Haplopolystichum, Dryopteridaceae): A new fern species from limestone area in Guizhou, China. Botanical Studies 50: 101-106 (cover article) Zhang L-B. 2004. Equisetaceae, Huperziaceae, Isoetaceae, Lycopodiaceae, Psilotaceae. Pp. 1-85, 224-245. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 6 (3), edited by Zhang X-C. Zhang L-B. 2001. Polystichum sect. Metapolystichum, sect. Neopolystichum. Pp. 52-60, 84-117. In: Wu C-Y (ed.), Flora Reipublicae Popularis Sinicae. Beijing: Science Press. Vol. 5 (2), edited by Kung H-S. Kung H-S and L-B Zhang. 2001. New pteridophyte records from Sichuan, China and their geographical significance. Fern Gazette 16: 195-199. Zhang L-B and H-S Kung. 2000a. The reclassification of Lycopodiaceae (s. str.) in China. Acta Phytotaxonomica Sinica 38: 266-275. Zhang L-B and H-S Kung. 2000b. An outline of the taxonomy and distribution of Equisetaceae in China. Pp. 82-88. In: Zhang X-C and K-H Shing (eds.), Ching Memorial Volume Beijing: China Forestry Publishing House. Zhang L-B and H-S Kung. 2000c. Taxonomy of Huperzia Bernh. (s. str.) sect. Serratae (Rothm.) Holub in China. Acta Phytotaxonomica Sinica 38: 13-22. Zhang L-B, and H-S Kung. 2000d. Two sections of Phlegmariurus (Herter) Holub (Huperziaceae) from China. Acta Phytotaxonomica Sinica 38: 23-29. Zhang L-B and H-S Kung. 1999a. On the taxonomy of Phlegmariurus (Herter) Holub (Huperziaceae) sect. Huperzioides H. S. Kung et L. B. Zhang (sect. nov.) in China. Acta Phytotaxonomica Sinica 37: 40-53. Zhang L-B and H-S Kung. 1999b. A new section of the fern genus Polystichum Roth ? sect. Neopolystichum Ching. Acta Phytotaxonomica Sinica 37: 81-86. Zhang L-B and H-S Kung. 1998a. A taxonomic study of Huperzia Berhn. (s. str.) sect. Huperzia in China. Acta Phytotaxonomica Sinica 36: 521-529. Zhang L-B and H-S Kung. 1998b. New materials of the fern genus Polystichum Roth from west China. Acta Phytotaxonomica Sinica 36: 465-468. Kung H-S and L-B Zhang. 1998. Study on the genus Polystichum Roth sect. Lasiopolystichum Daigobo from China. Acta Phytotaxonomica Sinica 36: 242-254. Kung H-S and L-B Zhang. 1996. The features of the pteridophyte flora of Mt. Gongga (Minya Gonkar), Sichuan, China. Acta Botanica Yunnanica 18: 176-184. Zhang L-B. 1996. On the foliar epidermis of Chinese sect. Metapolystichum Tagawa and sect. Neopolystichum (Polystichum, Dryopteridaceae) in China. Chinese Journal of Applied and Environmental Biology 2: 361-368. Zhang L-B and H-S Kung. 1996a. Taxonomic study on the fern genus Polystichum sect. Metapolystichum Tagawa from China (III). Acta Phytotaxonomica Sinica 34: 194-203. Zhang L-B and H-S Kung. 1996b. Taxonomic study on the fern genus Polystichum sect. Metapolystichum Tagawa from China (II). Acta Phytotaxonomica Sinica 34: 68-76. Zhang L-B and H-S Kung. 1995a. Taxonomic study on the fern genus Polystichum sect. Metapolystichum Tagawa from China (I). Acta Phytotaxonomica Sinica 33: 469-475. Zhang L-B and H-S Kung. 1995b. Four new species of the fern genus Polystichum sect. Metapolystichum from China. Acta Phytotaxonomica Sinica 33: 308-312. Kung H-S, L-B Zhang, and X-S Guo. 1995a. New materials of pteridophytes from Mt. Gongga (Minya Gonkar), Sichuan, China. Acta Botanica Yunnanica 17: 420-422. Kung H-S, L-B Zhang, and X-S Guo. 1995b. New materials of Dryopteris from Mt. Gongga (Minya Gonkar), Sichuan, China. Acta Botanica Yunnanica 17: 159-161. Zhang L-B, Z-R Wang, and X-S Guo. 1995a. New ferns of Athyriaceae from Mt. Gongga (Minya Gonkar), Sichuan, China. Acta Botanica Yunnanica 17: 302-304. Zhang L-B, H-S Kung, and X-S Guo. 1995b. New taxa of Polystichum Roth from Mt. Gongga (Minya Gonkar), Sichuan, China. Acta Botanica Yunnanica 17: 25-29. Zhang L-B and H-S Kung. 1994. Studies on the spore morphology of Chinese sect. Metapolystichum (Polystichum, Dryopteridaceae). Acta Botanica Yunnanica 16: 273-278. Kung H-S and L-B Zhang. 1994. A new species of Polystichum Roth from Burma and SW China. Acta Phytotaxnomica Sinica 32: 370-371. Zhang L-B. 1994. A taxonomic study on the fern genus Polystichum sect. Metapolystichum from Sichuan, China. Acta Botanica Yunnanica 16: 131-138. Zhang L-B. 1992. A study on the cladistic relationships of the genus Polystichum Roth sect. Metapolystichum Tagawa using the method of maximal same step length. Advances in Plant Taxonomy in Northern China, 1: 118-125.
IV. Systematics, phylogeny, and biogeography of flowering plants:
Liao M, LD Shepherd, J-Y Zhang, Y Feng, S Mattapha, L-B Zhang, X-F Gao, B Xu. 2023. Phylogeny, biogeography, and character evolution of the genus Sophora s.l. (Fabaceae, Papilionoideae). Molecular Phylogenetics and Evolution 181: 107713. Han M-Q, Yuan Q, Lu T-F, Zhang L-B & Liu Y. 2019. Petrocosmea qiruniae (Gesneriaceae), a new species from Guizhou, China with supplementary data and revised description of P. leiandra. Kew Bulletin 74(2): xx-xx. DOI: 10.1007/s12225-019-9811-8 Xu B, X-M Zeng, X-F Gao, P-J Dong & L-B Zhang. 2017. ITS non-concerted evolution and rampant hybridization in the legume genus Lespedeza (Fabaceae). Scientific Reports 7:40057. DOI: 10.1038/srep40057. Chen X, H He, and L-B Zhang. 2015. A monograph of the Anisophylleaceae (Cucurbitales) with description of 18 new species of Anisophyllea. Phytotaxa 229: 1–189. Xu B, Gao X-F & L-B Zhang. 2015. Lespedeza hengduanshanensis comb. & stat. nov. (Fabaceae) from the Hengduan Mountains of SW China. Phytotaxa 202: 165–168. Zhang L-B and MG Gilbert. 2015. Comparison of classifications of vascular plants of China. Taxon 64: 17–26. Le Péchon, T., Q. Dai, L-B Zhang, X-F. Gao, and H. Sauquet. 2015. Diversification of Dombeyoideae (Malvaceae) in the Mascarenes: Old taxa on young islands? International Journal of Plant Sciences 176: 211–221. Fougère-Danezan M, S Joly, A Bruneau, X-F Gao, and L-B Zhang. 2015. Phylogeny and biogeography of wild roses with specific attention to polyploids. Annals of Botany 115: 275–291. Chen X, H He, and L-B Zhang. 2013. A taxonomic revision of Anisophyllea (Anisophylleaceae) from Madagascar. Phytotaxa 148: 32–46. Wang, H-C, L-B Zhang, and Z-R He. 2013. Petrocosmea melanophthalma sp. nov. (sect. Deinanthera, Gesneriaceae), an attractive new species from Yunnan, China. Novon 22: 486–490. Xu, B., X.-F. Gao, and L-B Zhang. 2013. Lespedeza jiangxiensis, sp. nova. (Fabaceae) from China based on molecular and morphological evidence. Systematic Botany 38: 118–126. Zhang L-B and NT Lu. 2014. Solanum membranisepalum (Solanaceae): A new name for the Flora of China. Phytotaxa 186: 239–240. Zhang L-B. 2014. Callicarpa qianyiyongii (Lamiaceae): A new name for the Flora of China. Phytotaxa 178: 057–058. Duan Y-F and L-B Zhang. 2014. Eight new names and lectotypification of six names in Lamianae (asterids I) for the Flora of China. Phytotaxa 170: 278–282. Mao K, RI Milne, L-B Zhang, Y Peng, J Liu, P Thomas, RR Mill, and SS Renner. 2012. Distribution of living Cupressaceae reflects the breakup of Pangea. Proceedings of the National Academy of Sciences of the United States of America 109: 7793-7798. Su J-X, W Wang, L-B Zhang, and Z-D Chen. 2012. Phylogenetic placement of two enigmatic genera, Borthwickia and Stixis, based on molecular and pollen data, and the description of a new family of Brassicales, Borthwickiaceae. Taxon 61: 601-611. Xu B, N Wu, X-F Gao, and L-B Zhang. 2011. Molecular phylogeny of Lespedeza (Fabaceae) based on DNA sequences of six plastid and nuclear genes: Incongruence, introgression, and incomplete lineage sorting. Molecular Phylogenetics and Evolution 62: 346-358. Kadereit JW, H Goldner, N Holstein, G Schorr, and L-B Zhang. 2011. The stability of Quaternary speciation - a case study in Primula sect. Auricula. Alpine Botany DOI: 10.1007/s00035-010-0084-y. Meng J, M. Fougère-Danezan, L-B Zhang, D-Z Li, and T-S Yi. 2011. Untangling the hybrid origin of the Chinese tea rose: Evidence from DNA sequences of single-copy nuclear and chloroplast genes. Plant Systematics and Evolution 297: 157-170. Xu B, X-F Gao, N Wu, and L-B Zhang. 2011. Pollen diversity and its systematic implications in Lespedeza (Fabaceae). Systematic Botany 336: 352-361. Zhang G-C and L-B Zhang. 2011. Impatiens shimianensis sp. nov. (Balsaminaceae): A new species from Sichuan, China based on morphological and molecular evidence. Systematic Botany 36: 721-729. Michalak I, L-B Zhang, and SS Renner. 2010. Transatlantic, transpacific, and trans-Indian Ocean dispersal events in the small Gondwanan Laurales family Hernandiaceae. Journal of Biogeography 37: 1214-1226. Wei X-M, and X-F Gao, L-B Zhang. 2008. A systematic study of Rosa sericea (Rosaceae) complex: Are R. omeiensis and R. sericea conspecific? Journal of Systematics and Evolution 46: 919-928. Cusimano N, L-B Zhang, and SS Renner. 2008. Reevaluation of the cox1 group I intron in Araceae and angiosperms indicates a history dominated by loss rather than horizontal transfer. Molecular Biology and Evolution 25: 265-276. Zhang L-B, MP Simmons, and SS Renner. 2007. A phylogeny of Anisophylleaceae based on six nuclear and plastid loci: Ancient disjunctions and recent dispersal between South America, Africa, and Asia. Molecular Phylogenetics and Evolution. 44: 1057-1067. Kocyan A, L-B Zhang, H Schaefer, and SS Renner. 2007. A multi-locus chloroplast phylogeny for the Cucurbitaceae and its implications for character evolution and classification. Molecular Phylogenetics and Evolution. 44: 553-577. Zhang L-B and MP Simmons. 2006. Phylogeny and delimitation of the Celastrales inferred from nuclear and plastid genes. Systematic Botany 31: 107-121. Zhang L-B, MP Simmons, A Kocyan, and SS Renner. 2006. Phylogeny and evolution of the Cucurbitales based on DNA sequences of nine loci from three genomes. Molecular Phylogenetics and Evolution 39: 305-322. Zhang L-B and JW Kadereit. 2004. Classification of Primula sect. Auricula (Primulaceae) based on two molecular data sets (ITS, AFLPs), morphology and geographical distribution. Botanical Journal of the Linnean Society 146: 1-26. Renner SS, L-B Zhang, and J Murata. 2004. A chloroplast phylogeny of Arisaema (Araceae) illustrates Tertiary floristic links between Asia, North America, and East Africa. American Journal of Botany 91: 881-888. Renner SS and L-B Zhang. 2004. Biogeography of the Pistia clade (Araceae) based on cp and mtDNA sequences and Bayesian divergence time inference. Systematic Biology 53: 422-432. Zhang L-B, HP Comes, and JW Kadereit. 2004. The temporal course of Quaternary diversification in the European high mountain endemic Primula sect. Auricula (Primulaceae). International Journal of Plant Sciences 165: 191-207 (cover article). Zhang L-B and SS Renner. 2003. The deepest splits in Chloranthaceae as resolved by chloroplast sequences. International Journal of Plant Sciences 164(6 Suppl.): 383-392. Zhang L-B and JW Kadereit. 2002. The systematics of Soldanella L. (Primulaceae) based on morphological and molecular (ITS, AFLPs) evidence. Nordic Journal of Botany 22: 129-169. Zhang L-B. 2002. Phylogeny, biogeography, and systematics of Soldanella L. and Primula L. sect. Auricula Duby (Primulaceae) based on molecular and morphological evidence. Goettingen: Cuvillier Verlag. 157pp. Zhang L-B, HP Comes, and JW Kadereit. 2001. Phylogeny and Quaternary history of the European montane/alpine endemic Soldanella (Primulaceae) based on ITS and AFLP variation. American Journal of Botany 88: 2331-2345.
V. Theoretical systematics:
Simmons MP, L-B Zhang, and K Müller. 2010. Phylogenetic inference using non-redundant coding of dependent characters vs. alternative approaches when applied to protein-coding genes. Cladistics 26: 1-11. Simmons MP, L-B Zhang, CT Webb, and K Mueller. 2007. A penalty of using anonymous dominant markers (AFLPs, ISSRs, and RAPDs) for phylogenetic inference. Molecular Phylogenetics and Evolution 42: 528-542. Simmons MP, L-B Zhang, CT Webb, and A Reeves. 2006. How can third codon positions outperform first and second codon positions in phylogenetic inference? An empirical example from the seed plants. Systematic Biology 55: 245-258. Simmons MP*, L-B Zhang*, CT Webb, A Reeves, and JA Miller. 2006. The relative performance of Bayesian and parsimony approaches when sampling characters evolving under homogeneous and heterogeneous sets of parameters. Cladistics 22: 171-185 (*co-first authors).
VI. Botanical nomenclature:
Zhou X-M, X Zhang & L-B Zhang. 2018. Proposal to reject the name Selaginella flagellifera (Selaginellaceae). Taxon 67: 641–642. Rothfels, CR, Costa J, Zhang L-B, Moran RC, Salino A, Smith AR. 2018. (2579) Proposal to conserve Aspidium draconopterum (Draconopteris draconoptera) (Tectariaceae) with a conserved type. Taxon 67: 204–205. Lu TN, X-F Gao & L-B Zhang. 2017. Proposal to conserve the name Polystichum omeiense against P. caruifolium (Dryopteridaceae). Taxon 66: 1468–1469. Peng Y-L & L-B Zhang. 2016. Comments on author citations for nomenclatural novelties published in volume 15 of the Flora Reipublicae Popularis Sinicae (FRPS). Phytotaxa 269: 236. Ji C-F and L-B Zhang. 2016. Lectotypification of 11 names of Eurya (Pentaphylacaceae) for flora of China. Phytotaxa 269: 61–64. Duan Y-F and L-B Zhang. 2014. Proposal to conserve the name Ligustrum sempervirens (Franch.) Lingelsh. against L. sempervirens Lindl. (Oleaceae). Taxon 63: 945–946. Zhang L-B. 2012. The nomenclature of the lycophyte species Phlegmariurus mingcheensis Ching (Huperziaceae). Taxon 61: 665-666. Zhang L-B. 2011. [Chinese Translation of] Knapp S, McNeill J, and NJ Turland. 2011. Changes to publication requirements made at the XVIII International Botanical Congress in Melbourne - what does e-publication mean for you? PhytoKey 6: 13-19 & Plant Systematics & Resources 33: 514-517. Zhang L-B. 2007. [Chinese Translation of] McNeill, J, FR Barrie, HM Burdet, V Demoulin, DL Hawksworth, K Marhold, DH Nicolson, J Prado, PC Silva, JE Skog, JH Wiersema, and NJ Turland (eds.). 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress Vienna, Austria, July 2005. Beijing: Science Press & St. Louis: Missouri Botanical Garden Press. Zhang L-B, Q-E Yang, NJ Turland, and J McNeill. 2007. Main changes in the new edition of the International Code of Botanical Nomenclature - the "Vienna Code". Acta Phytotaxonomica Sinica 45 (2): 251-255. Zhang L-B, Silva, PC, J McNeill, and NJ Turland. 2007. On the terms in the International Code of Botanical Nomenclature. Acta Phytotaxonomica Sinica 45: 593-598. Zhang L-B and JW Kadereit. 2005. Typification and synonymization in Primula sect. Auricula (Primulaceae). Taxon 54: 775-788. Zhang L-B and JW Kadereit. 2004a. Nomenclature of Soldanella L. (Primulaceae). Taxon 53: 741-752.